Castilleja miniata var. fulva

(Pennell) J. M. Egger

Phytologia 90: 72. 2008.

Endemic
Basionyms: Castilleja fulva Pennell Proc. Acad. Nat. Sci. Philadelphia 86: 540. 1934
Treatment appears in FNA Volume 17. Treatment on page 629. Mentioned on page 579, 628.

Herbs 4–5 dm; with a taproot. Stems glabrous or glabrate proximally, hairy distally, hairs sparse, spreading, long, soft to ± stiff, short-stipitate-glandular. Leaves narrowly lanceolate to narrowly ovate, thin and not fleshy, margins plane, flat, usually 0-lobed, apex acuminate to acute. Bracts distally pale yellowish, pale orange, or salmon, sometimes pale-red, 0–3 (–7) -lobed, central lobe sometimes distally apiculate; central lobe apex rounded, lateral ones acute to acuminate. Calyces 15–21 mm; abaxial and adaxial clefts 10 mm, 50–70% of calyx length, abaxial slightly deeper than adaxial, lateral 1–6 mm, 5–30% of calyx length; lobes narrowly lanceolate to narrowly triangular, apex acute to acuminate. Corollas 20–27 mm; beak 9–11 mm; abaxial lip green, 1–1.5 mm, 10–20% as long as beak. 2n = 96.


Phenology: Flowering Jun–Aug.
Habitat: Damp low thickets, grassy verges, roadsides, forest edges.
Elevation: 600–1600 m.

Discussion

Variety fulva is found widely in central and northern British Columbia, as well as in the Peace River valley in adjacent west-central Alberta and southern Yukon, substantially but not entirely replacing var. miniata in these regions. The inflorescences are consistently yellow to pale tawny orange within populations and have a distinctive morphology, though overlapped by the wide range of variation in var. miniata. It is not known if the distinctive coloration of var. fulva represents introgression from Castilleja unalaschcensis, or if these populations might have diverged in isolation on a nunatak during the last extensive glaciation, as postulated by F. W. Pennell (1934b).

Selected References

None.

Lower Taxa

None.

"thin" is not a number."medium" is not a number.

0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
ascending +  and incurved +
not inflated +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Acute (?) +  and Acuminate (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(Pennell) J. M. Egger +
Castilleja fulva +
0.9 cm9 mm <br />0.009 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
not +, leathery +  and fleshy +
pale-red;salmon;pale orange;salmon;pale orange;pale yellowish +
0-3(-7)-lobed +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
2-lobed +, lobed +  and tubular +
1.5 cm15 mm <br />0.015 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
0-5(-7)-lobed +, lanceolate +  and oblong-ovate +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
2 cm20 mm <br />0.02 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
Alta. +, B.C. +  and Yukon. +
600–1600 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Damp low thickets, grassy verges, roadsides, forest edges. +
much-branched +, unbranched +  and eglandular +
short-stipitate-glandular +
short to long +
15 cm150 mm <br />0.15 m <br /> (22 cm220 mm <br />0.22 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
8 cm80 mm <br />0.08 m <br /> (9.5 cm95 mm <br />0.095 m <br />) +
green +  and purple +
deciduous +
basal +  and cauline +
narrowly lanceolate;narrowly ovate +
3 cm30 mm <br />0.03 m <br /> (8 cm80 mm <br />0.08 m <br />) +
not fleshy +
above mid lobes +, arising +, erect +  and ascending-spreading +
3 +, 2 +  and 4 +
narrowly lanceolate +  and narrowly triangular +
subequal +
0-lobed +  and plane +
tenuinucellate +  and unitegmic +
campylotropous-like +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering Jun–Aug. +
slender +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
non-secund +  and secund +
elongate +
subequal +
branched +  and rooting +
creeping +  and decumbent +
aerial +  and subterranean +
erect +  and ascending +
not fleshy +
2-lobed +  and entire +
Euchroma +  and Oncorhynchus +
Castilleja miniata var. fulva +
Castilleja miniata +
variety +
white +  and green +
shallow;deep +
erect +  and incurved +
reduced +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +