Grimmia mollis

Bruch & Schimper

Bryol. Europ. 3: 133, plate 253. 1849,.

Synonyms: Hydrogrimmia mollis (Bruch & Schimper) I. Hagen Grimmia evansii E. Britton
Treatment appears in FNA Volume 27. Treatment on page 257. Mentioned on page 226.

Plants in loose patches, green. Stems 1–5 cm, central strand present. Leaves oblong to ovatelanceolate, 2–3 × 0.8–1 mm, margins plane, incurved distally, muticous, concave, apex rounded to cucullate, costa not projecting on abaxial side, subpercurrent; basal juxtacostal laminal cells quadrate, thin-walled; basal marginal laminal cells quadrate, thin-walled; medial laminal cells quadrate, thin-walled; juxtacostal and marginal distal laminal cells 1-stratose. Sexual condition dioicous. Seta straight, 2–3 mm, yellowish. Capsule sporadically present, exserted, yellowish, ovoid to oblong, smooth, exothecial cells thick-walled, annulus absent, operculum rostellate, peristome teeth purple, perforated, fully developed, papillose. Calyptra cucullate.


Habitat: Wet acidic rocks in alpine and boreal habitats
Elevation: moderate to high elevations (1000-4100 m)

Distribution

V27 358-distribution-map.gif

Greenland, Alta., B.C., Que., Yukon, Alaska, Calif., Colo., Mont., Wash., Eurasia

Discussion

Grimmia mollis, discovered by Schimper in the Tirol of Austria, is a characteristic arctic-alpine species, frequently growing in glacial streams, sometimes over reaches of hundreds of meters. Although the species is widespread in the Northern Hemisphere, it is nowhere common. It occurs exclusively above the treeline. Because of its unique subpercurrent costa, uniform lamina of quadrate, thin-walled cells, and peculiar ecological niche, it cannot easily be confused with any other species. I. Hagen (1909) placed it in a monotypic genus, Hydrogrimmia. This concept was followed by M. F. V. Corley et al. (1981). However, there are no significant morphological characters to separate it from Grimmia, prompting H. C. Greven (2003) to follow, in agreement with H. A. Crum and L. E. Anderson (1981), Bruch and Schimper’s original concept. This dioicous species grows in separate male and female cushions, and capsules are produced very rarely.

Selected References

None.

Lower Taxa

None.
... more about "Grimmia mollis"
deciduous +  and persistent +
well-defined +
differentiated +
Roxanne I. Hastings +  and Henk C. Greven +
Bruch & Schimper +
only rarely longer +
not papillose +
toothed +
sinuous +  and straight to sometimes +
sinuose-nodulose;quadrate;elongate +
oblate;elongate +
divided +
undifferentiated +
plicate +  and smooth +
less +  and 1/2 +
not papillose +
cucullate +
small;medium +
sulcate +  and smooth +
pendent +  and erect +
striate +  and smooth +
ovoid;oblong +
thin +  and thick +
thick-walled +  and thin-walled +
quadrate +  and rectangular +
excurrent;percurrent;excurrent +
not projecting +
subpercurrent +
reniform +, elliptical +  and terete +
usually not markedly larger +
1-stratose +  and multistratose +
Greenland +, Alta. +, B.C. +, Que. +, Yukon +, Alaska +, Calif. +, Colo. +, Mont. +, Wash. +  and Eurasia +
moderate to high elevations (1000-4100 m) +
reproduction +  and asexual +
dioicous +, autoicous +  and sexual +
Wet acidic rocks in alpine and boreal habitats +
papillose +, mammillose +  and smooth +
specialized +
dioicous +, autoicous +  and sexual +
2mm;3mm +
oblong +  and ovatelanceolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
1;multistratose +
concave +, muticous +  and plane +
thick-walled +  and sinuose +
thick-walled +  and sinuose +
rostellate +
not +  and enlarged +
not +  and differentiated +
lanceolate;linear +
Bryol. Europ. +
straight +
sigmoid +  and arcuate +
papillose +  and smooth +
short to long +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
ornamented +  and smooth +
globose +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
muticous;long-awned +
Hydrogrimmia mollis +  and Grimmia evansii +
Grimmia mollis +
Grimmia subg. Undetermined +
species +