Heterotheca villosa var. nana

(A. Gray) Semple

Novon 4: 54. 1994.

Endemic
Basionyms: Chrysopsis canescens var. nana A. Gray Mem. Amer. Acad. Sci., n. s. 4: 78. 1849
Synonyms: Chrysopsis horrida Rydberg Heterotheca horrida (Rydberg) V. L. Harms Heterotheca villosa var. horrida (Rydberg) Semple
Treatment appears in FNA Volume 20. Treatment on page 253. Mentioned on page 250, 251, 254.

Stems ascending to erect (1–60+), 9–40 (–63) cm, moderately to densely hispido-strigose, abundantly long-hirsute, sparsely to densely stipitate-glandular. Distal cauline leaf-blades (often perpendicluar to stems), linear to broadly oblong, (6–) 10–24 (–32) mm, ± reduced distally, bases rounded, margins flat, apices usually acute, often obtuse, faces moderately hispido-strigose (hairs 9–56/mm2), moderately to densely stipitate-glandular (glands 2–36/mm2). Heads 2–20 (–33) in usually congested, corymbiform arrays. Peduncles (4–) 10–35 (–48) mm, moderately to densely hispido-strigose, moderately to densely glandular; bracts 1–4, linear-oblong, reduced distally, not leaflike. Involucres cylindric to campanulate (fresh), (5.4–) 6.4–8.4 (–9) mm. Phyllaries narrowly triangular-lanceolate, margins sometimes reddish purple distally, faces sparsely to moderately strigose, sparsely to densely glandular. Ray-florets 11–19 (–22); laminae (3.5–) 6.9–11 (–12) mm. 2n = 18.


Phenology: Flowering (May–)Jun–Aug(–Oct).
Habitat: Sandy and gravelly limestone, sandstone and granitic soils, fissures in lava surfaces, igneous cliffs, short-grass grasslands, dry sagebrush areas, open mixed conifer forests at lower elevations, pinyon-juniper-cholla brush
Elevation: 1300–2700(–2900) m

Distribution

V20-552-distribution-map.gif

Ariz., Colo., Kans., Nebr., N.Mex., Tex., Utah, Wyo.

Discussion

Variety nana is distinguished by its linear to broadly oblong, often perpendicular, distal stem leaves and often congested, corymbiform arrays. Forms of var. minor with oblong to ovate leaves generally have stems with longer internodes and fewer leaves. Plants at higher elevations in the foothills approach narrow-leaved forms of Heterotheca fulcrata in general appearance, and may hybridize with them, especially in the Laramie Hills of Wyoming. Possible hybrids between var. nana and var. foliosa occur in Colorado and Wyoming, where the two are sympatric. In the Four Corners area, var. nana grades into var. scabra, and collections often are difficult to place into one of the two taxa; both have broader leaf bases than var. minor as it occurs in the region. Rare plants in Wisconsin show similarities to both var. minor and var. nana. The variety was long known under the epithet horrida (either as a species or variety), but at the varietal rank the epithet nana is older.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
paniculiform +, corymbiform +  and open +
indeterminate +  and determinate +
John C. Semple +
(A. Gray) Semple +
rounded +
Chrysopsis canescens var. nana +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
linear-oblong +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
2.4 cm24 mm <br />0.024 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
linear +  and broadly oblong +
Ariz. +, Colo. +, Kans. +, Nebr. +, N.Mex. +, Tex. +, Utah +  and Wyo. +
1300–2700(–2900) m +
strigose +, moderately hispido-strigose +  and moderately densely stipitate-glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Sandy and gravelly limestone, sandstone and granitic soils, fissures in lava surfaces, igneous cliffs, short-grass grasslands, dry sagebrush areas, open mixed conifer forests at lower elevations, pinyon-juniper-cholla brush +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.84 cm8.4 mm <br />0.0084 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
cylindric;campanulate +
0.64 cm6.4 mm <br />0.0064 m <br /> (0.84 cm8.4 mm <br />0.0084 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.69 cm6.9 mm <br />0.0069 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
3.5 cm35 mm <br />0.035 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
hispido-strigose +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
Flowering (May–)Jun–Aug(–Oct). +
triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
1 (?) +  and 60 (?) +
ascending +  and erect +
1 +  and 50 +
9 cm90 mm <br />0.09 m <br /> (40 cm400 mm <br />0.4 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis horrida +, Heterotheca horrida +  and Heterotheca villosa var. horrida +
Heterotheca villosa var. nana +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +