Pinus torreyana

Parry ex Carrière

326. 1855.

Common names: Torrey pine
EndemicConservation concern
Treatment appears in FNA Volume 2.

Trees to 15 (–23) m; trunk to 1m diam., in nature mostly crooked and leaning; crown rounded to flattened or irregular. Bark redbrown to purple-red, deeply furrowed with irregular, elongate, flat, scaly ridges. Branches irregular, spreading-ascending, candelabralike; twigs stout (1–2cm thick), greenish, aging deep gray-brown to near black, rough. Buds conic-ovoid, pale-brown, to 2.5cm; scale margins white-fringed. Leaves mostly 5 per fascicle, ascending or spreading, persisting 3–4 years, 15–30cm × ca. 2mm, straight or curved, slightly twisted, dull gray-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly acute; sheath to 2cm, shed early, base persistent. Pollen cones ovoid, 20–30mm, yellow. Seed-cones maturing in 3 years, shedding seeds soon thereafter, persisting to 5 years, lateral, massive, heavy, symmetric, ovoid before opening, broadly ovoid when open, 10–15cm, yellow to redbrown, lustrous, stalks to 4cm; apophyses thick, angulately dome-shaped, with 5 low convergent keels; umbo central, forming short, curved-tipped pyramid. Seeds narrowly obovoid; body 16–24mm, brown, apically dark-brown; wing broad, oblique-tipped, to 15mm.

Discussion

Pinus torreyana is a rare and local Tertiary relic species whose present range is reduced to two small areas of southern California: near Del Mar (San Diego County) and on the northeastern shore of Santa Rosa Island (Santa Barbara County). Its distribution in Oligocene and Miocene (or at least that of its near ancestor) extended north to Oregon. Its harsh natural habitat elicits an unusually contorted and often sparse form, quite unlike the cleaner and taller form the species takes in cultivation.

In terms of numbers of individuals in the wild, as well as the small area occupied by natural populations, Pinus torreyana is without a doubt the rarest North American pine. As such it is under protection. Artificial crosses between it and another, more widespread Tertiary relic, P. sabiniana, have been successful.

Subspecies 2.

Key

1 Crown of sheltered tree narrower than height of mature tree, open with well-spaced branches in natural sites; seed cones less than 13.5 cm broad, as broad as long or longer; only tip of umbo curved outward; maximum seed width ca. 12 mm. Pinus torreyana subsp. torreyana
1 Crown of sheltered tree broader than height of mature tree, compact in natural sites; seed cones mostly more than 13.5 cm broad, broader than long; entire umbo curved outward; maximum seed width ca. 14 mm. Pinus torreyana subsp. insularis
... more about "Pinus torreyana"
resinous +
dome--shaped +
Robert Kral +
Parry ex Carrière +
redbrown;purple-red +
dark-brown +  and brown +
1.6 cm16 mm <br />0.016 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
spreading-ascending +
candelabra-like +
pale-brown +
conspicuous +
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Torrey pine +
yellow;redbrown +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
rounded +  and flattened +
Calif. +  and only in the flora +
sheathed +  and sessile +
rounded +, 2-3-angled +  and terete +
arranged +  and fascicles +
gray-green +
curved +  and straight +
persisting +
15cm;30cm +
3 (?) +  and 4 (?) +
scale-like +
2 cm20 mm <br />0.02 m <br /> (?) +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not winged +  and 2-winged +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
curved-tipped +
haller1986a +
elongate +
fibrous;woody +
arranged +  and overlapping +
persistent +
white-fringed +
flattened +
pliable +  and woody +
relatively thin;strongly thickened +
grouped +  and solitary +
long-persistent +
wingless +, winged +, stalked +  and sessile +
persistent +
pendent;more or less erect +
obovoid +
persistent +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
reduced;well-defined short +
Endemic +  and Conservation concern +
4 cm40 mm <br />0.04 m <br /> (?) +
extended +
Apinus +, Strobus +  and Caryopitys +
Pinus torreyana +
species +
decurrent +  and elongate +
gray-brown +  and greenish +
oblique-tipped +
reduced +
0 cm0 mm <br />0 m <br /> (?) +  and 1.5 cm15 mm <br />0.015 m <br /> (?) +
5 +  and 3 +