Sorbus sitchensis var. grayi

(Wenzig) C. L. Hitchcock in C. L. Hitchcock et al.

in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. 3: 189. 1961.

Common names: Dwarf or western mountain ashE
Endemic
Basionym: Sorbus sambucifolia var. grayi Wenzig Bot. Centralbl. 35: 342. 1888
Synonyms: Pyrus occidentalis S. Watson S. occidentalis (S. Watson) Greene S. sitchensis subsp. grayi (Wenzig) Calder & Roy L. Taylor
Treatment appears in FNA Volume 9. Treatment on page 444. Mentioned on page 445.

Leaflets: l/w ratio 1.9–3.5, margins entire or toothed apically, serrate for less than 1/2 their length, serrations fine to coarse. Flowers: petals white or pinkish, white or pink in bud, claw villous; styles (2–) 2.4–3.2 mm. 2n = 34.


Phenology: Flowering spring (late summer); fruiting fall.
Habitat: Mountain slopes, forest edges, rockslides, edges of avalanche tracks, peatlands, muskegs, meadows
Elevation: 0–3000 m

Distribution

V9 750-distribution-map.jpg

B.C., Alaska, Calif., Oreg., Wash.

Discussion

Variety grayi often has smaller leaflets than var. sitchensis, and plants near timberline frequently are dwarfed by exposure. It is usually found at higher elevations or on boggier soils than var. sitchensis. The rank is controversial; it was treated by G. N. Jones (1939) and H. A. McAllister (2005) as a species (Sorbus occidentalis). The only reliable character to segregate it is the extent of toothing on the leaf margins, which is variable on some individuals, with nearly entire leaflets on shoots exposed to sun and with leaflet margins to 50% toothed on shaded basal shoots.

Variety grayi occurs south to Crater Lake National Park in Klamath County, Oregon (P. F. Zika 2003). Most collections and reports from California (T. J. Rosatti 1993) were misidentified, except for a single sheet from Siskiyou County in the Klamath Mountains (R. White 207, HSC). One of the synonyms, Pyrus occidentalis, was described with several syntypes from California, which are not var. grayi but Sorbus californica. G. N. Jones (1939, p. 40) effectively lectotypified P. occidentalis with a syntype from the Canadian border (Lyall s.n., GH), which has nearly entire leaflets and is var. grayi.

Selected References

None.

Lower Taxa

None.
Peter F. Zika +  and Stéphane M. Bailleul +
(Wenzig) C. L. Hitchcock in C. L. Hitchcock et al. +
grayish red +, initially reddish purple +  and brownish gray +
compound +  and simple +
opposite +  and alternate +
not +  and clawed +
Sorbus sambucifolia var. grayi +
blue-green +  and paler +
shiny +  and dull +
ovate +  and oblong +
5 cm50 mm <br />0.05 m <br /> (33 cm330 mm <br />0.33 m <br />) +
? (?) +  and ? (?) +
not glutinous +
red-purple +  and redbrown +
villous +  and glaucous +
0.8 cm8 mm <br />0.008 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
odd +  and simple +
divided +  and lobed +
tanniferous +
Dwarf or western mountain ashE +
B.C. +, Alaska +, Calif. +, Oreg. +  and Wash. +
not +  and aggregated +
0–3000 m +
single-layered +
1.1 cm11 mm <br />0.011 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
rancid +, unpleasant +  and strong +
unisexual +  and bisexual +
not +  and aggregated +
not +  and aggregated +
yellow +, brown +, red +  and orange +
glabrous +, hairy +  and glaucous +
0.4 cm4 mm <br />0.004 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
Mountain slopes, forest edges, rockslides, edges of avalanche tracks, peatlands, muskegs, meadows +
whitish +  and rufous +
green +  and red +
glaucous +  and glabrous +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
villous +, glaucous +  and glabrous +
crassinucellate +
deciduous +
subopposite +  and opposite +
0.6 cm6 mm <br />0.006 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
papillose +
obovate +, ovate +, oblanceolate +, oblong +  and narrowly elliptic +
serrate +, entire or +  and finely coarsely serrate +
toothed +  and entire +
toothed +, unlobed +  and lobed +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
3 +  and 2 +
12-80-flowered +
2 cm20 mm <br />0.02 m <br /> (8 cm80 mm <br />0.08 m <br />) +
hairy +, glaucous +  and glabrous +
glaucous or +  and sparsely densely villous +
glaucous or +  and sparsely densely villous +
0 (?) +  and 4 (?) +
Pink (?) +, White (?) +  and Pinkish (?) +
free +  and distinct +
2 +  and 5 +
0.4 cm4 mm <br />0.004 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
Flowering spring (late summer) +  and fruiting fall. +
adnate +, free +, connate +  and distinct +
red +  and pinkish red +
0.7 cm7 mm <br />0.007 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
subglobose +  and broadly elliptic or obovoid +
0.7 cm7 mm <br />0.007 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
in C. L. Hitchcock et al., Vasc. Pl. Pacif. N.W. +
not arillate +
0.34 cm3.4 mm <br />0.0034 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
flattened;trigonous;symmetric;ovoid;ovoid-lanceolate oblong or elliptic +
0.22 cm2.2 mm <br />0.0022 m <br /> (?) +  and 0.29 cm2.9 mm <br />0.0029 m <br /> (?) +
persistent +
free +  and distinct +
inconspicuous +
triangular +  and ovate +
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
free +  and distinct +
branched +  and simple +
ascending +  and erect +
1 +  and 8 +
deciduous +  and persistent +
short-adnate +  and free +
0.2 cm2 mm <br />0.002 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
elongate +
0.24 cm2.4 mm <br />0.0024 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
Pyrus occidentalis +, S. occidentalis +  and S. sitchensis subsp. grayi +
Sorbus sitchensis var. grayi +
Sorbus sitchensis +
variety +
inconspicuous +
hairy +  and glabrous +