View source for Achlys ← Achlys You do not have permission to edit this page, for the following reason: The action you have requested is limited to users in the group: Users. You can view and copy the source of this page. {{Treatment/ID |accepted_name=Achlys |accepted_authority=de Candolle |publications={{Treatment/Publication |title=Syst. Nat. |place=2: 35. 1821 |year=1821 }} |basionyms= |synonyms= |hierarchy=Berberidaceae;Achlys |hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Berberidaceae]]</div><div class="higher-taxon"><small>genus</small>[[Achlys]]</div></div> |etymology=Greek Achlus, a god of night |volume=Volume 3 |mention_page= |treatment_page= }}<!-- --><span class="statement" id="st-d0_s0" data-properties="whole_organism duration;whole_organism duration;whole_organism some measurement;whole_organism pubescence;whole_organism growth form"><b>Herbs,</b> perennial, deciduous, 2.5-5 dm, glabrous.</span> <span class="statement" id="st-d0_s1" data-properties="rhizome size;rhizome architecture;foliage leaf quantity;shoot life cycle"><b>Rhizomes </b>extensive, branching, producing 1-few foliage leaves or flowering shoots per year.</span> <span class="statement" id="st-d0_s2" data-properties="stem location;stem presence"><b>Aerial </b>stems absent.</span> <span class="statement" id="st-d0_s3" data-properties="leaf position;leaf arrangement;leaf architecture"><b>Leaves </b>basal, alternate, 3-foliolate;</span> <span class="statement" id="st-d0_s4" data-properties="petiole length or size;petiole size">petiole long, slender.</span> <span class="statement" id="st-d0_s5" data-properties="blade-leaf shape;blade-leaf distance"><b>Leaf-</b>blade orbiculate in gross outline, 20-40 cm;</span> <span class="statement" id="st-d0_s6" data-properties="leaflet blade shape;leaflet blade shape;leaflet blade shape;leaflet blade architecture or shape">leaflet blades fan-shaped, entire or lobed, lateral leaflet blades strongly asymmetric, margins entire to coarsely sinuate;</span> <span class="statement" id="st-d0_s7" data-properties="margin shape;margin architecture">venation palmate.</span> <span class="statement" id="st-d0_s8" data-properties="inflorescence position or structure subtype;spike density;spike architecture;flower prominence"><b>Inflorescences </b>terminal, dense scapose-pedunculate spikes of inconspicuous flowers.</span> <span class="statement" id="st-d0_s9" data-properties="flower architecture;flower coloration;flower some measurement;flower some measurement"><b>Flowers </b>3-merous, white to cream, 6 mm or less;</span> <span class="statement" id="st-d0_s10" data-properties="bracteole presence">bracteoles absent;</span> <span class="statement" id="st-d0_s11" data-properties="sepal presence">sepals absent;</span> <span class="statement" id="st-d0_s12" data-properties="petal presence">petals absent;</span> <span class="statement" id="st-d0_s13" data-properties="stamen atypical quantity;stamen quantity">stamens 8-10 (-13);</span> <span class="statement" id="st-d0_s14" data-properties="anther dehiscence;flap quantity">anthers dehiscing by 2 apically hinged flaps;</span> <span class="statement" id="st-d0_s15" data-properties="pollen exine coloration or pubescence or relief">pollen exine striate;</span> <span class="statement" id="st-d0_s16" data-properties="">ovaries asymmetrically ellipsoid;</span> <span class="statement" id="st-d0_s17" data-properties="ovary shape;ovary placentation;placenta development">placentation marginal, placenta developed only near base of ovary.</span> <span class="statement" id="st-d0_s18" data-properties="fruits follicle coloration;fruits follicle coloration;fruits follicle course;fruits follicle architecture"><b>Fruits </b>follicles with transverse dehiscence, purplish red or brown, curved, furrowed.</span> <span class="statement" id="st-d0_s19" data-properties="seed quantity;seed coloration"><b>Seeds </b>1, brown;</span> <span class="statement" id="st-d0_s20" data-properties="">aril absent.</span> <span class="statement" id="st-d0_s21" data-properties="aril presence;x chromosome quantity">x = 6.</span><!-- -->{{Treatment/Body |distribution=North America;Asia (Japan) |discussion=<p>Species 3 (2 in the flora).</p><!-- --><p>Achlys is of particular interest because of its amphi-Pacific disjunction. Despite the 8000 km or more disjunction, the taxa are remarkably similar in morphology, ecology, and karyology. Japanese populations are diploid; American populations are diploid and tetraploid.</p><!-- --><p>Two species are recognized in this treatment; some researchers prefer to treat them as varieties. In the Californian portion of the range, some field botanists believe the two taxa are sufficiently morphologically distinct to be called species; farther north these differences are reportedly less apparent.</p> |tables= |references= }}<!-- --><div class="treatment-key"> ==Key== <div class="treatment-key-group"> {| class="wikitable fna-keytable" |-id=key-0-1 |1 |Central leaflet blade 4–11 cm, distal margins (1–)3–4(–8)-lobed; stamens 3–4 mm; follicles red-purple. |[[Achlys triphylla|Achlys triphylla]] |-id=key-0-1 |1 |Central leaflet blade ca. 7–16 cm, distal margins (3–)6–9(–12)-lobed; stamens 3.8–6 mm; follicles brown. |[[Achlys californica|Achlys californica]] |} </div></div><!-- -->{{#Taxon: name=Achlys |author=R. David Whetstone;T.A. Atkinson;Daniel D. Spaulding |authority=de Candolle |rank=genus |parent rank=family |synonyms= |basionyms= |family=Berberidaceae |distribution=North America;Asia (Japan) |reference=None |publication title=Syst. Nat. |publication year=1821 |special status= |source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V3/V3_251.xml |genus=Achlys |anther dehiscence=dehiscing |aril presence=absent |blade-leaf distance=20cm;40cm |blade-leaf shape=orbiculate |bracteole presence=absent |flap quantity=2 |flower architecture=3-merous |flower coloration=white;cream |flower prominence=inconspicuous |flower some measurement=less;6 |foliage leaf quantity=1;few |fruits follicle architecture=furrowed |fruits follicle coloration=brown;purplish red |fruits follicle course=curved |inflorescence position or structure subtype=terminal |leaf architecture=3-foliolate |leaf arrangement=alternate |leaf position=basal |leaflet blade architecture or shape=asymmetric |leaflet blade shape=lobed;entire;fan--shaped |margin architecture=palmate |margin shape=entire;coarsely sinuate |ovary placentation=marginal |ovary shape=ellipsoid |petal presence=absent |petiole length or size=long |petiole size=slender |placenta development=developed |pollen exine coloration or pubescence or relief=striate |rhizome architecture=branching |rhizome size=extensive |seed coloration=brown |seed quantity=1 |sepal presence=absent |shoot life cycle=flowering |spike architecture=scapose-pedunculate |spike density=dense |stamen atypical quantity=10;13 |stamen quantity=8;10 |stem location=aerial |stem presence=absent |whole_organism duration=deciduous;perennial |whole_organism growth form=herb |whole_organism pubescence=glabrous |whole_organism some measurement=2.5dm;5dm |x chromosome quantity=6 }}<!-- -->[[Category:Treatment]][[Category:Berberidaceae]] Templates used on this page: Template:Berberidaceae (view source) Template:Treatment/AuthorLink (view source) Template:Treatment/Body (view source) Template:Treatment/Body/Maps (view source) Template:Treatment/ID (view source) Template:Treatment/Publication (view source) Return to Achlys.