View source for Anomodon

{{Treatment/ID
|accepted_name=Anomodon
|accepted_authority=Hooker & Taylor
|publications={{Treatment/Publication
|title=Muscol. Brit.,
|place=79, plates 3 [near upper right], 33 [upper center left & right]. 1818
|year=1818
}}
|basionyms=
|synonyms={{Treatment/ID/Synonym
|name=Haplohymenium
|authority=Dozy & Molkenboer
|rank=genus
}}
|hierarchy=Anomodontaceae;Anomodon
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Anomodontaceae]]</div><div class="higher-taxon"><small>genus</small>[[Anomodon]]</div></div>
|etymology=Greek, anomalos, abnormal, and odon, tooth, alluding to reduced peristome
|volume=Volume 28
|mention_page=page 341, 342, 630, 637, 652
|treatment_page=page 629
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism pubescence;whole_organism coloration;whole_organism growth form"><b>Plants </b>small to large, ± glaucous, green to rusty brown.</span> <span class="statement" id="st-d0_s1" data-properties="stem course or shape;stem shape;stem architecture;branch orientation"><b>Stems </b>with branches erect to arcuate, sometimes attenuate to flagellate distally;</span> <span class="statement" id="st-d0_s2" data-properties="strand cell variability;strand cell variability">central strand cells differentiated or not;</span> <span class="statement" id="st-d0_s3" data-properties="pseudoparaphyllium presence;species quantity">pseudoparaphyllia absent in all but 2 species.</span> <span class="statement" id="st-d0_s4" data-properties="branch leaf fixation or orientation;branch leaf architecture;branch leaf shape;branch leaf shape;branch leaf orientation;branch leaf arrangement;branch leaf shape;mid leaf shape;mid leaf architecture or arrangement or shape or vernation"><b>Branch </b>leaves appressed to moderately secund or crispate when dry, complanate or erect to imbricate when moist, broadly ovate to lanceolate, ± abruptly narrowed mid leaf, not plicate proximally (somewhat plicate in A. longifolius);</span> <span class="statement" id="st-d0_s5" data-properties="margin shape;margin shape;margin shape;margin architecture or shape;margin relief;margin shape;margin architecture or shape;margin shape">margins plane, sometimes undulate or revolute, entire, papillose, crenulate, serrulate, or sometimes denticulate near apex;</span> <span class="statement" id="st-d0_s6" data-properties="apex shape;apex shape;apex shape;apex shape;apex shape;apex shape">apex rounded, obtuse, acute, or narrowly acuminate;</span> <span class="statement" id="st-d0_s7" data-properties="costa fragility;costa prominence;costa shape;costa cell relief;costa cell relief">costa usually strong, sometimes obscured by laminal cells and ending before mid leaf, sometimes discreetly, asymmetrically 2-fid at end, abaxial costa cells smooth or papillose;</span> <span class="statement" id="st-d0_s8" data-properties="laminal cell shape;laminal cell prominence;laminal cell architecture or course;laminal cell size;papilla quantity;papilla quantity;papilla height;wall width">laminal cells hexagonal, obscure to irregular, small, papillae 1 or many, high, on both surfaces, walls thin;</span> <span class="statement" id="st-d0_s9" data-properties="basal-cell shape;basal-cell coloration;basal-cell architecture or pubescence or relief;wall size">basal-cells sometimes oblong, pellucid, smooth, walls incrassate.</span> <span class="statement" id="st-d0_s10" data-properties="perichaetial leaf variability"><b>Perichaetial </b>leaves well differentiated.</span> <span class="statement" id="st-d0_s11" data-properties="seta some measurement"><b>Seta </b>to 2.2 cm.</span> <span class="statement" id="st-d0_s12" data-properties="capsule architecture or shape"><b>Capsule </b>symmetric;</span> <span class="statement" id="st-d0_s13" data-properties="stoma presence">stomata sometimes present;</span> <span class="statement" id="st-d0_s14" data-properties="peristome size">peristome reduced (well developed in A. rostratus);</span> <span class="statement" id="st-d0_s15" data-properties="exostome tooth coloration;exostome tooth relief;exostome tooth pubescence;exostome tooth architecture">exostome teeth white to pale-brown, densely papillose, occasionally cross-striolate, sometimes slightly trabeculate;</span> <span class="statement" id="st-d0_s16" data-properties="endostome size;endostome presence;basal membrane quantity;cell height;segment shape;segment size;segment presence">endostome sometimes very reduced or absent (sect. Haplohymenium), basal membrane 2–7 cells high, segments keeled to linear and reduced or absent.</span> <span class="statement" id="st-d0_s17" data-properties="calyptra relief;calyptra pubescence"><b>Calyptra </b>smooth to papillose or hirsute (sect. Haplohymenium).</span> <span class="statement" id="st-d0_s18" data-properties="spore atypical some measurement;spore some measurement"><b>Spores </b>9–20 (–23) µm.</span><!--

-->{{Treatment/Body
|distribution=North America;Mexico;West Indies;Central America;South America (Bolivia);Europe;s Africa;Pacific Islands (New Zealand);Australia;temperate;circumboreal regions
|discussion=<p>Species 16 (8 in the flora).</p><!--
--><p>Anomodon viticulosus grows mostly on shaded calcareous outcrops, but the other species are found on tree trunks, including their base, logs, or sometimes soil or rock. Anomodon attenuatus, A. minor, and A. rostratus may grow on the same tree, with A. rugelii sometimes joining them in submontane regions. Although A. tristis may be found growing with the above species, it usually forms much thinner, more delicate mats higher on the tree. In North America, at least two species (A. attenuatus and A. rostratus) fruit profusely; A. minor and A. rugelii fruit less abundantly and perhaps less frequently, while sporophytes of A. viticulosus are extremely rare in North America (only one fertile specimen of A. viticulosus seen, none of A. tristis).</p><!--
--><p>Haplohymenium was created to accommodate plants that resemble Anomodon but are more slender and have a papillose calyptra with long, hyaline scattered hairs. Segregating Haplohymenium would make the rest of Anomodon paraphyletic, as Haplohymenium is a sister group of A. minor, A. rugelii, and A. viticulosus, all of which are part of subg. Anomodon (Í. Granzow-de la Cerda 1997); these taxa are more distantly related to subg. Pseudoanomodon (Limpricht) Ochyra, to which A. attenuatus and A. rostratus belong.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Branch leaves long-lanceolate; apices ending in hair-point or subulalike
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Branch leaves ligulate; apices rounded, obtuse, acute, or apiculate, not ending in hair-point or subulalike
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Stems profusely branched; primary branches erect.
|[[Anomodon rostratus|Anomodon rostratus]]
|-id=key-0-2
|2
|Stems sparingly branched; primary branches prostrate or pendulous.
|[[Anomodon longifolius|Anomodon longifolius]]
|-id=key-0-3
|3
|Plants small; stems less than 1 mm thick when dry; branch leaves less than 2.1 mm; apices often broken off; costa ending much before apex, obscured by laminal cells distally
|[[#key-0-4| > 4]]
|-id=key-0-3
|3
|Plants small to large; stems usually more than 0.8 mm thick when dry; branch leaves sometimes greater than 2 mm; apices intact; costa ending near apex, not or rarely obscured by laminal cells distally
|[[#key-0-5| > 5]]
|-id=key-0-4
|4
|Stems 0.3-0.5 mm thick when dry; leaves 0.5-0.9 mm; basal laminal cells few, region not reaching margin; costa weak, obscured by laminal cells almost throughout.
|[[Anomodon tristis|Anomodon tristis]]
|-id=key-0-4
|4
|Stems 0.5-1 mm thick when dry; leaves 1.2-1.8(-2.1) mm; basal laminal cells many, region reaching margin; costa moderately strong, obscured by laminal cells distally.
|[[Anomodon thraustus|Anomodon thraustus]]
|-id=key-0-5
|5
|Plants dark green to rusty brown; leaves incurved-contorted when dry; bases auriculate; costae golden yellow to rusty brown; pseudoparaphyllia present.
|[[Anomodon rugelii|Anomodon rugelii]]
|-id=key-0-5
|5
|Plants green to yellowish; leaves erect, imbricate, appressed or rarely slightly crisped when dry; bases broadly decurrent; costae pellucid or light green; pseudoparaphyllia absent
|[[#key-0-6| > 6]]
|-id=key-0-6
|6
|Plants large; stems 1-1.8 mm thick when dry; leaves flexuose, secund, spreading when moist, erect when dry, greater than 2 mm.
|[[Anomodon viticulosus|Anomodon viticulosus]]
|-id=key-0-6
|6
|Plants medium-sized to large; stems usually less than 1 mm thick when dry; leaves complanate when moist, appressed when dry, usually less than 2 mm
|[[#key-0-7| > 7]]
|-id=key-0-7
|7
|Stems not pinnate, secondary branches not attenuate, often slightly clavate at apices; perichaetia on terminal branches, beyond distalmost branching points; leaves abruptly narrowed mid leaf; apices rounded; margins entire at apex; abaxial costa cells with rounded-simple papillae in rows.
|[[Anomodon minor|Anomodon minor]]
|-id=key-0-7
|7
|Stems irregularly pinnate, secondary branches attenuate at apices; perichaetia never present beyond distalmost branching points; leaves slightly narrowed mid leaf; apices acute, sometimes obtuse or slightly apiculate; margins sometimes denticulate at apex; abaxial costa cells smooth.
|[[Anomodon attenuatus|Anomodon attenuatus]]
|}
</div></div><!--

-->{{#Taxon:
name=Anomodon
|authority=Hooker & Taylor
|rank=genus
|parent rank=family
|synonyms=Haplohymenium
|basionyms=
|family=Anomodontaceae
|distribution=North America;Mexico;West Indies;Central America;South America (Bolivia);Europe;s Africa;Pacific Islands (New Zealand);Australia;temperate;circumboreal regions
|reference=None
|publication title=Muscol. Brit.,
|publication year=1818
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_988.xml
|genus=Anomodon
|apex shape=acuminate;acute;acuminate;acute;obtuse;rounded
|basal membrane quantity=2;7
|basal-cell architecture or pubescence or relief=smooth
|basal-cell coloration=pellucid
|basal-cell shape=oblong
|branch leaf architecture=secund
|branch leaf arrangement=imbricate
|branch leaf fixation or orientation=appressed
|branch leaf orientation=erect
|branch leaf shape=broadly ovate;lanceolate
|branch orientation=erect
|calyptra pubescence=hirsute
|calyptra relief=smooth;papillose
|capsule architecture or shape=symmetric
|cell height=high
|costa cell relief=papillose;smooth
|costa fragility=strong
|costa prominence=obscured
|costa shape=2-fid
|endostome presence=absent
|endostome size=reduced
|exostome tooth architecture=trabeculate
|exostome tooth coloration=white;pale-brown
|exostome tooth pubescence=cross-striolate
|exostome tooth relief=papillose
|laminal cell architecture or course=irregular
|laminal cell prominence=obscure
|laminal cell shape=hexagonal
|laminal cell size=small
|margin architecture or shape=serrulate;entire
|margin relief=papillose
|margin shape=denticulate;crenulate;revolute;undulate;plane
|mid leaf architecture or arrangement or shape or vernation=plicate
|mid leaf shape=narrowed
|papilla height=high
|papilla quantity=many;1
|perichaetial leaf variability=differentiated
|peristome size=reduced
|pseudoparaphyllium presence=absent
|segment presence=absent
|segment shape=keeled;linear
|segment size=reduced
|seta some measurement=0cm;2.2cm
|species quantity=2
|spore atypical some measurement=20um;23um
|spore some measurement=9um;20um
|stem architecture=flagellate
|stem course or shape=arcuate
|stem shape=attenuate
|stoma presence=absent
|strand cell variability=not;differentiated
|wall size=incrassate
|wall width=thin
|whole_organism coloration=green;rusty brown
|whole_organism growth form=plant
|whole_organism pubescence=glaucous
|whole_organism size=small;large
}}<!--

-->[[Category:Treatment]][[Category:Anomodontaceae]]