View source for Bryhnia

{{Treatment/ID
|accepted_name=Bryhnia
|accepted_authority=Kaurin
|publications={{Treatment/Publication
|title=Bot. Not.
|place=1892: [60]. 1892
|year=1892
}}
|basionyms=
|synonyms=
|hierarchy=Brachytheciaceae;Bryhnia
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Brachytheciaceae]]</div><div class="higher-taxon"><small>genus</small>[[Bryhnia]]</div></div>
|etymology=For Nils Bryhn, 1854 – 1916, Norwegian bryologist
|volume=Volume 28
|mention_page=page 405, 407, 408, 429, 447, 461, 652
|treatment_page=page 428
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism coloration;whole_organism growth form;tuft architecture or fragility;tuft density"><b>Plants </b>small to medium-sized, in moderately loose to dense tufts, green or yellowish to brownish.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem shape;stem architecture;stem architecture or shape;stem architecture or shape;stem architecture or shape;branch shape;branch architecture;branch architecture or shape"><b>Stems </b>creeping to ascending, not attenuate, unevenly foliate, julaceous or not, irregularly to regularly pinnate, branches moderately densely terete to complanate-foliate, sometimes julaceous;</span> <span class="statement" id="st-d0_s2" data-properties="central strand presence">central strand present;</span> <span class="statement" id="st-d0_s3" data-properties="pseudoparaphyllium shape">pseudoparaphyllia acute;</span> <span class="statement" id="st-d0_s4" data-properties="">axillary hairs of 3 or 4 cells.</span> <span class="statement" id="st-d0_s5" data-properties="stem-leaf orientation;stem-leaf orientation;stem-leaf orientation;stem-leaf orientation;stem-leaf orientation;stem-leaf fixation or orientation;stem-leaf shape;stem-leaf shape;stem-leaf shape;stem-leaf shape;stem-leaf architecture or arrangement or shape or vernation"><b>Stem-</b>leaves erect, patent, or rigidly spreading, imbricate-appressed, ovate, ovate-triangular, or lanceolate, moderately to strongly concave, not or slightly plicate;</span> <span class="statement" id="st-d0_s6" data-properties="base shape">base decurrent;</span> <span class="statement" id="st-d0_s7" data-properties="margin architecture or shape;margin architecture or shape">margins serrulate proximally, serrate distally;</span> <span class="statement" id="st-d0_s8" data-properties="apex shape;apex shape;apex shape;apex architecture or shape;apex shape;apex shape;apex shape;apex shape">apex gradually tapered, acute, acuminate, truncate, apiculate, or cucullate;</span> <span class="statement" id="st-d0_s9" data-properties="leaf character;leaf width">costa to 40–80% leaf length, broad throughout, terminal abaxial spine present or absent;</span> <span class="statement" id="st-d0_s10" data-properties="terminal abaxial spine presence;terminal abaxial spine presence;cell shape;cell size">alar cells subquadrate to short-rectangular, large;</span> <span class="statement" id="st-d0_s11" data-properties="laminal cell course;wall width">laminal cells elongate-flexuose, walls moderately to strongly thick, prorate;</span> <span class="statement" id="st-d0_s12" data-properties="basal-cell height or length or size;basal-cell width">basal-cells shorter, wider.</span> <span class="statement" id="st-d0_s13" data-properties="branch leaf size;branch leaf width"><b>Branch </b>leaves smaller, narrower;</span> <span class="statement" id="st-d0_s14" data-properties="apex shape">apex acute to acuminate (sharper than stem-leaves);</span> <span class="statement" id="st-d0_s15" data-properties="costal abaxial surface architecture or shape">costal abaxial surface more strongly serrate;</span> <span class="statement" id="st-d0_s16" data-properties="">laminal cells more strongly prorate.</span> <span class="statement" id="st-d0_s17" data-properties="laminal cell reproduction;laminal cell reproduction"><b>Sexual </b>condition dioicous;</span> <span class="statement" id="st-d0_s18" data-properties="leaf acumen orientation">perichaetial leaf acumen reflexed.</span> <span class="statement" id="st-d0_s19" data-properties="seta coloration;seta pubescence or relief"><b>Seta </b>brownish orange to redbrown, rough.</span> <span class="statement" id="st-d0_s20" data-properties="capsule orientation;capsule coloration;capsule shape;capsule course"><b>Capsule </b>inclined to horizontal, brownish orange to redbrown, cylindric, not or slightly curved;</span> <span class="statement" id="st-d0_s21" data-properties="annulus arrangement">annulus separating by fragments;</span> <span class="statement" id="st-d0_s22" data-properties="operculum shape;operculum shape">operculum long-conic, broadly rostrate;</span> <span class="statement" id="st-d0_s23" data-properties="peristome architecture;peristome architecture">peristome xerocastique, perfect.</span> <span class="statement" id="st-d0_s24" data-properties="calyptra architecture"><b>Calyptra </b>naked.</span> <span class="statement" id="st-d0_s25" data-properties="spore some measurement"><b>Spores </b>13–18 µm.</span><!--

-->{{Treatment/Body
|distribution=North America;Eurasia
|discussion=<p>Species 5–7 (3 in the flora).</p><!--
--><p>The circumscription of Bryhnia needs a re-evaluation with DNA markers since morphology seems to be misleading. In Japan and adjacent areas, N. Takaki (1956) accepted 15 species within Bryhnia, but only six survived the revision by A. Noguchi and Z. Iwatsuki (1987+); most were synonymized with B. novae-angliae. Among North American species, B. graminicolor is not closely related to the core group and possibly should be segregated in its own genus.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Plants small; stems to 2 cm; stem leaves narrowly ovate to lanceolate, 0.6-1.1 × 0.2-0.5 mm.
|[[Bryhnia graminicolor|Bryhnia graminicolor]]
|-id=key-0-1
|1
|Plants medium-sized; stems 2.5-10 cm; stem leaves broadly ovate-triangular to ovate, 0.7-1.3(-1.6) × 0.7-1.2 mm
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Stem leaves ovate-triangular to ovate, 1-1.3(-1.6) × 0.7-1.1 mm, longer than broad; apices acute or acuminate, rarely cucullate; stems rarely julaceous along some portions; alar regions gradually differentiated; e North America.
|[[Bryhnia novae-angliae|Bryhnia novae-angliae]]
|-id=key-0-2
|2
|Stem leaves broadly ovate-triangular, 0.7-1.2 × 0.8-1.2 mm, often shorter than broad; apices broadly acute to rounded-truncate and short-apiculate, often cucullate; stems julaceous; alar regions abruptly differentiated; Alaska, British Columbia.
|[[Bryhnia hultenii|Bryhnia hultenii]]
|}
</div></div><!--

-->{{#Taxon:
name=Bryhnia
|author=Michael S. Ignatov
|authority=Kaurin
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Brachytheciaceae
|distribution=North America;Eurasia
|reference=None
|publication title=Bot. Not.
|publication year=1892
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_666.xml
|genus=Bryhnia
|annulus arrangement=separating
|apex architecture or shape=truncate
|apex shape=acute;acuminate
|basal-cell height or length or size=shorter
|basal-cell width=wider
|base shape=decurrent
|branch architecture=complanate-foliate
|branch architecture or shape=julaceous
|branch leaf size=smaller
|branch leaf width=narrower
|branch shape=terete
|calyptra architecture=naked
|capsule coloration=brownish orange;redbrown
|capsule course=curved
|capsule orientation=inclined;horizontal
|capsule shape=cylindric
|cell shape=subquadrate;short-rectangular
|cell size=large
|central strand presence=absent
|costal abaxial surface architecture or shape=serrate
|laminal cell course=elongate-flexuose
|laminal cell reproduction=dioicous;sexual
|leaf acumen orientation=reflexed
|leaf character=length
|leaf width=broad
|margin architecture or shape=serrate;serrulate
|operculum shape=rostrate;long-conic
|peristome architecture=perfect;xerocastique
|pseudoparaphyllium shape=acute
|seta coloration=brownish orange;redbrown
|seta pubescence or relief=rough
|spore some measurement=13um;18um
|stem architecture=foliate
|stem architecture or shape=pinnate;not;julaceous
|stem orientation=creeping;ascending
|stem shape=not attenuate
|stem-leaf architecture or arrangement or shape or vernation=plicate
|stem-leaf fixation or orientation=imbricate-appressed
|stem-leaf orientation=spreading;patent;spreading;patent;erect
|stem-leaf shape=lanceolate;moderately strongly concave
|terminal abaxial spine presence=absent;absent
|tuft architecture or fragility=loose
|tuft density=dense
|wall width=thick
|whole_organism coloration=yellowish;brownish
|whole_organism growth form=plant
|whole_organism size=small;medium-sized
}}<!--

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