View source for Campylium

{{Treatment/ID
|accepted_name=Campylium
|accepted_authority=(Sullivant) Mitten
|publications={{Treatment/Publication
|title=J. Linn. Soc., Bot.
|place=12: 631. 1869
|year=1869
}}
|basionyms={{Treatment/ID/Basionym
|name=Hypnum sect. Campylium
|authority=Sullivant
|rank=section
|publication_title=in A. Gray, Manual ed.
|publication_place=2, 677. 1856
}}
|synonyms=
|hierarchy=Amblystegiaceae;Campylium
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Amblystegiaceae]]</div><div class="higher-taxon"><small>genus</small>[[Campylium]]</div></div>
|etymology=Greek campylos, curved, alluding to reflexed leaves
|volume=Volume 28
|mention_page=page 265, 288, 290, 291, 292, 295, 305, 306, 315, 529, 549, 560, 645
|treatment_page=page 287
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism size;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism growth form"><b>Plants </b>small or medium-sized, green, yellowish green, golden brown, golden yellow, or brownish.</span> <span class="statement" id="st-d0_s1" data-properties="stem architecture;stem architecture;stem architecture;stem architecture;stem architecture"><b>Stems </b>unbranched, irregularly branched, or irregularly pinnate;</span> <span class="statement" id="st-d0_s2" data-properties="hyalodermi presence;central strand presence">hyalodermis absent, central strand absent;</span> <span class="statement" id="st-d0_s3" data-properties="paraphyllium presence;paraphyllium presence">paraphyllia absent or sometimes present;</span> <span class="statement" id="st-d0_s4" data-properties="rhizoid insertion;rhizoid relief;rhizoid relief;rhizoid initial insertion;rhizoid initial relief;rhizoid initial relief">rhizoids or rhizoid initials on stem or abaxial costa insertion, rarely forming tomentum, slightly or strongly branched, smooth or slightly warty-papillose;</span> <span class="statement" id="st-d0_s5" data-properties="distal cell quantity;distal cell coloration">axillary hair distal cells 1–5, hyaline.</span> <span class="statement" id="st-d0_s6" data-properties="leaf-stem orientation;leaf-stem orientation;leaf-stem orientation;leaf-stem orientation;leaf-stem orientation;leaf-stem arrangement;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem architecture or arrangement or shape or vernation;leaf-stem distance"><b>Stem-</b>leaves erect, spreading, or ± squarrose, occasionally distinctly falcate-secund, cordate, cordate-ovate, ovate, or rounded-triangular, not plicate, 1–4.6 mm;</span> <span class="statement" id="st-d0_s7" data-properties="base shape">base not or hardly decurrent;</span> <span class="statement" id="st-d0_s8" data-properties="margin shape;margin shape;margin shape;limbidium presence">margins plane, entire or slightly sinuate, limbidia absent;</span> <span class="statement" id="st-d0_s9" data-properties="apex shape;acumen variability;acumen variability;acumen architecture">apex acuminate, acumen differentiated or not, furrowed;</span> <span class="statement" id="st-d0_s10" data-properties="leaf position;leaf quantity">costa double or single, ending below mid leaf;</span> <span class="statement" id="st-d0_s11" data-properties="cell variability;cell shape;cell shape;cell shape;cell shape;cell shape;cell coloration;widest cell width;region fusion;region shape;region shape;region shape;region shape;region shape;region shape">alar cells differentiated, rectangular, quadrate, or sometimes transversely rectangular or short-rectangular, strongly inflated, hyaline, widest cells 17–30 µm wide, region distinct, quadrate, broadly ovate, ovate, or rectangular, along basal margins, reaching from margin 15–60% distance to costa at insertion;</span> <span class="statement" id="st-d0_s12" data-properties="medial laminal cell arrangement or course or shape">medial laminal cells linear, rarely shortly so;</span> <span class="statement" id="st-d0_s13" data-properties="">marginal cells 1-stratose.</span> <span class="statement" id="st-d0_s14" data-properties="marginal cell architecture;marginal cell reproduction;marginal cell reproduction;marginal cell reproduction"><b>Sexual </b>condition dioicous or autoicous.</span> <span class="statement" id="st-d0_s15" data-properties="capsule orientation;capsule shape;capsule course"><b>Capsule </b>horizontal, cylindric, curved;</span> <span class="statement" id="st-d0_s16" data-properties="peristome architecture">peristome perfect;</span> <span class="statement" id="st-d0_s17" data-properties="exostome margin architecture or shape;exostome margin architecture or shape">exostome margins dentate or slightly dentate distally;</span> <span class="statement" id="st-d0_s18" data-properties="endostome cilium quantity;endostome cilium development;endostome cilium shape">endostome cilia usually 1–3, well developed, nodulose.</span> <span class="statement" id="st-d0_s19" data-properties="spore atypical some measurement;spore some measurement"><b>Spores </b>(10–) 11–24 µm.</span><!--

-->{{Treatment/Body
|distribution=North America;Mexico;West Indies (Haiti);Central America (Guatemala);Eurasia;Pacific Islands (New Zealand)
|discussion=<p>Species 4 (4 in the flora).</p><!--
--><p>Campylium occurs in more or less mineral-rich wetlands that are mostly permanently wet or moist. Differences between members of this genus and those of 6. Campyliadelphus and 18. Campylophyllum are discussed with the latter two genera. Members of Drepanocladus with similarly oriented leaves have transversely triangular alar groups that extend from the margin 60–100% of the distance to the costa at the insertion. Other species of Drepanocladus may have approximately quadrate alar groups, but not ovate or rectangular and extending up the leaf margin as in Campylium. The typification of Campylium was discussed by P. Isoviita and L. Hedenäs (1997).</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Sexual condition autoicous; stem leaves erect or spreading; bases subsheathing, broadly cordate-ovate, ovate, or narrowly ovate; acumina when differentiated constituting at most 33% leaf length
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Sexual condition dioicous; stem leaves spreading or ± squarrose; bases erect to erect-spreading, cordate or rounded-triangular; acumina frequently differentiated, constituting (33-)40-77% leaf length
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Stem leaves slightly concave, 1.8-2.4 mm; bases ± broadly cordate-ovate; leaves gradually narrowed to apex; acumina not differentiated.
|[[Campylium laxifolium|Campylium laxifolium]]
|-id=key-0-2
|2
|Stem leaves concave or strongly so, 2.1-4.6 mm; bases ovate or narrowly ovate; leaves ± suddenly or more gradually narrowed to apex; acumina frequently differentiated.
|[[Campylium longicuspis|Campylium longicuspis]]
|-id=key-0-3
|3
|Stems erect, irregularly branched or sometimes irregularly pinnate; stem leaves 1.7-2.8 × 0.7-1.2 mm; acumina constituting 40-65% (rarely 33-40% in Arctic plants) leaf length; paraphyllia absent.
|[[Campylium stellatum|Campylium stellatum]]
|-id=key-0-3
|3
|Stems creeping, irregularly pinnate; stem leaves 1-2.3 × 0.4-1 mm; acumina constituting 55-77% leaf length; paraphyllia sometimes present.
|[[Campylium protensum|Campylium protensum]]
|}
</div></div><!--

-->{{#Taxon:
name=Campylium
|author=Lars Hedenäs
|authority=(Sullivant) Mitten
|rank=genus
|parent rank=family
|synonyms=
|basionyms=Hypnum sect. Campylium
|family=Amblystegiaceae
|distribution=North America;Mexico;West Indies (Haiti);Central America (Guatemala);Eurasia;Pacific Islands (New Zealand)
|reference=None
|publication title=J. Linn. Soc., Bot.
|publication year=1869
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_448.xml
|genus=Campylium
|acumen architecture=furrowed
|acumen variability=not;differentiated
|apex shape=acuminate
|base shape=decurrent
|capsule course=curved
|capsule orientation=horizontal
|capsule shape=cylindric
|cell coloration=hyaline
|cell shape=inflated;short-rectangular;rectangular;quadrate;rectangular
|cell variability=differentiated
|central strand presence=absent
|distal cell coloration=hyaline
|distal cell quantity=1;5
|endostome cilium development=developed
|endostome cilium quantity=1;3
|endostome cilium shape=nodulose
|exostome margin architecture or shape=dentate;dentate
|hyalodermi presence=absent
|leaf position=mid
|leaf quantity=single
|leaf-stem architecture or arrangement or shape or vernation=not plicate
|leaf-stem arrangement=falcate-secund
|leaf-stem distance=1mm;4.6mm
|leaf-stem orientation=squarrose;spreading;squarrose;spreading;erect
|leaf-stem shape=rounded-triangular;ovate;rounded-triangular;ovate;cordate-ovate;cordate
|limbidium presence=absent
|margin shape=sinuate;entire;plane
|marginal cell architecture=1-stratose
|marginal cell reproduction=autoicous;dioicous;sexual
|medial laminal cell arrangement or course or shape=linear
|paraphyllium presence=absent;absent
|peristome architecture=perfect
|region fusion=distinct
|region shape=rectangular;ovate;rectangular;ovate;ovate;quadrate
|rhizoid initial insertion=branched
|rhizoid initial relief=warty-papillose;smooth
|rhizoid insertion=branched
|rhizoid relief=warty-papillose;smooth
|spore atypical some measurement=10um;11um
|spore some measurement=11um;24um
|stem architecture=pinnate;branched;pinnate;branched;unbranched
|whole_organism coloration=brownish;golden yellow;brownish;golden yellow;golden brown;yellowish green;green
|whole_organism growth form=plant
|whole_organism size=medium-sized;small
|widest cell width=17um;30um
}}<!--

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