View source for Ceratodon ← Ceratodon You do not have permission to edit this page, for the following reason: The action you have requested is limited to users in the group: Users. You can view and copy the source of this page. {{Treatment/ID |accepted_name=Ceratodon |accepted_authority=Bridel |publications={{Treatment/Publication |title=Bryol. Univ. |place=1: 480. 1826 , }} |basionyms= |synonyms= |hierarchy=Ditrichaceae;Ceratodon |hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Ditrichaceae]]</div><div class="higher-taxon"><small>genus</small>[[Ceratodon]]</div></div> |etymology=Greek keratos, horn, and odon, tooth, alluding to peristome teeth forked like goat horns |volume=Volume 27 |mention_page=page 443, 444, 458 |treatment_page=page 445 }}<!-- --><span class="statement" id="st-d0_s0" data-properties="whole_organism architecture or fragility;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism growth form;tuft density;turf density;mat coloration"><b>Plants </b>in loose to dense tufts, turfs, or mats, green to dark green, brownish green, light green, or yellow-green, often tinged reddish-brown or purple.</span> <span class="statement" id="st-d0_s1" data-properties="stem atypical some measurement;stem atypical some measurement;stem some measurement;stem architecture"><b>Stems </b>(0.2–) 1–3 (–4) cm, often branched;</span> <span class="statement" id="st-d0_s2" data-properties="rhizoid relief">rhizoids at base, papillose.</span> <span class="statement" id="st-d0_s3" data-properties="leaf orientation;leaf arrangement or shape;leaf shape;leaf course;leaf shape;leaf shape;leaf shape;leaf shape;leaf shape;margin orientation;margin shape;margin architecture;margin architecture"><b>Leaves </b>erect-patent, contorted or somewhat crisped, rarely straight when dry, lanceolate, ovatelanceolate, or triangular-lanceolate, or elliptical to ovate and somewhat concave, margins recurved to near apex, rarely plane, irregularly serrate or smooth distally;</span> <span class="statement" id="st-d0_s4" data-properties="costa position;costa architecture;awn length or size;awn architecture or pubescence or relief;awn quantity;row quantity">costa subpercurrent to excurrent, sometimes as a long, smooth awn, 1 row of guide cells, two stereid bands, hydroid cells present between guide cells and abaxial stereids;</span> <span class="statement" id="st-d0_s5" data-properties="leaf size">medial laminal cells of somewhat uneven shape and size across leaf, more or less quadrate or shortrectangular, often irregularly angled or rarely rounded, non-pitted.</span> <span class="statement" id="st-d0_s6" data-properties="propagule reproduction;propagule presence;propagule architecture;propagule texture;wall width;wall arrangement"><b>Specialized </b>asexual reproduction absent or as multicellular filamentous propagules with thin walls scattered along the stems or occasionally as rhizome nodules.</span> <span class="statement" id="st-d0_s7" data-properties="medial laminal cell size;medial laminal cell shape;medial laminal cell shape;medial laminal cell relief;medial laminal cell development;medial laminal cell reproduction;medial laminal cell reproduction"><b>Sexual </b>condition dioicous;</span> <span class="statement" id="st-d0_s8" data-properties="whole_organism reproduction;whole_organism reproduction;whole_organism growth form">male and female plants about the same size;</span> <span class="statement" id="st-d0_s9" data-properties="perigonial leaf shape;perigonial leaf shape;perigonial leaf shape;perigonial leaf shape;plant development">perigonial leaves ovate, concave, short-acuminate or rarely long-acuminate in well-developed plants;</span> <span class="statement" id="st-d0_s10" data-properties="perichaetial leaf shape;perichaetial leaf shape">perichaetial leaves convolute-sheathing, abruptly subulate to gradually acuminate.</span> <span class="statement" id="st-d0_s11" data-properties="seta coloration;seta coloration;seta coloration;seta coloration;seta coloration;seta coloration;seta shape;seta architecture;seta orientation"><b>Seta </b>red, purplish, yellow, or yellow-orange, elongate, twisted when dry, erect.</span> <span class="statement" id="st-d0_s12" data-properties="capsule orientation;capsule position;capsule coloration;capsule coloration;capsule coloration;capsule shape;capsule architecture or shape;capsule architecture;capsule architecture;capsule shape"><b>Capsule </b>erect to inclined or horizontal, exserted, dark red to reddish or purplish brown, to pale-brown, pale-yellow or yellow-orange, oblong-ovoid to oblong-cylindric or cylindric, often somewhat asymmetric and deeply furrowed, smooth to strongly sulcate when dry, usually strumose;</span> <span class="statement" id="st-d0_s13" data-properties="row quantity;cell size;cell duration;cell architecture">annulus of 2–3 rows of large, deciduous, revoluble cells;</span> <span class="statement" id="st-d0_s14" data-properties="operculum shape;operculum course">operculum conic to long-conic, straight;</span> <span class="statement" id="st-d0_s15" data-properties="peristome quantity;tooth quantity;segment quantity;segment shape;segment fusion;node quantity;basal membrane presence;basal membrane relief">peristome single, teeth 16, split nearly to their base into 2 filiform segments free to united at their nodes basally, 5–18 articulations, basal membrane present, finely papillose to spinulose-papillose.</span> <span class="statement" id="st-d0_s16" data-properties="calyptra shape"><b>Calyptra </b>cucullate.</span> <span class="statement" id="st-d0_s17" data-properties="spore shape;spore relief"><b>Spores </b>globose, smooth to finely papillose.</span><!-- -->{{Treatment/Body |distribution=Worldwide |discussion=<p>Species 3 (2 in the flora).</p><!-- --><p>Historically, Ceratodon has been a troubling genus. Taxonomic interpretations, especially with respect to C. purpureus in the broad sense, have varied widely, mainly because of the high degree of environmental and suspected genetic variation across its range. J. S. Burley and N. M. Pritchard (1990) provided the most thorough treatment of Ceratodon to date, reducing the number of species to four and subspecies of C. purpureus to three. One of their species, C. conicus, is treated here as a subspecies of C. purpureus, based on the apparent gradation and reduction of all of the characters that they used in their treatment. However, there remains a great need for a detailed study of this genus within North America.</p> |tables= |references={{Treatment/Reference |id=burley1990a |text=Burley, J. S. and N. M. Pritchard. 1990. Revision of the genus Ceratodon (Bryophyta). Harvard Pap. Bot. 2: 17–76. }}{{Treatment/Reference |id=snider1994a |text=Snider, J. A. 1994. Ceratodon. In: A. J. Sharp, et al., eds. The moss flora of Mexico. Mem. New York Bot. Gard. 69: 103. }} }}<!-- --><div class="treatment-key"> ==Key== <div class="treatment-key-group"> {| class="wikitable fna-keytable" |-id=key-0-1 |1 |Vegetative leaves elliptic to ovate-lanceolate, concave, obtuse; distal laminal cells usually 12-16 mm, sometimes longer; leaf margins plane to weakly recurved and usually entire; most costa sub-percurrent; capsule ovate to ovate-cylindrical, about 1 mm; spores usually19-21 µm; restricted to Arctic regions. |[[Ceratodon heterophyllus|Ceratodon heterophyllus]] |-id=key-0-1 |1 |Vegetative leaves usually lanceolate to ovate-lanceolate, acute to short-acuminate; distal laminal cells usually 8-12 mm; leaf margins recurved and usually toothed distally; costa percurrent to long-excurrent; capsule ovate-cylindrical to cylindrical, usually longer, to 3 mm; spores usually 11-14 µm; widespread. |[[Ceratodon purpureus|Ceratodon purpureus]] |} </div></div><!-- -->{{#Taxon: name=Ceratodon |author=Terry T. McIntosh |authority=Bridel |rank=genus |parent rank=family |synonyms= |basionyms= |family=Ditrichaceae |distribution=Worldwide |reference=burley1990a;snider1994a |publication title=Bryol. Univ. |publication year= |special status= |source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27_640.xml |genus=Ceratodon |awn architecture or pubescence or relief=smooth |awn length or size=long |awn quantity=1 |basal membrane presence=absent |basal membrane relief=finely papillose;spinulose-papillose |calyptra shape=cucullate |capsule architecture=smooth;strongly sulcate |capsule architecture or shape=asymmetric |capsule coloration=dark red;reddish or purplish brown |capsule orientation=erect;inclined or horizontal |capsule position=exserted |capsule shape=strumose;oblong-ovoid;oblong-cylindric or cylindric |cell architecture=revoluble |cell duration=deciduous |cell size=large |costa architecture=excurrent |costa position=subpercurrent |leaf arrangement or shape=contorted |leaf course=straight |leaf orientation=erect-patent |leaf shape=concave;triangular-lanceolate or elliptical;ovate |leaf size=quadrate |margin architecture=smooth;serrate |margin orientation=recurved |margin shape=plane |mat coloration=green;dark green brownish green light green or yellow-green |medial laminal cell development=specialized |medial laminal cell relief=non-pitted |medial laminal cell reproduction=dioicous;sexual |medial laminal cell shape=rounded;angled |medial laminal cell size=shortrectangular |node quantity=5;18 |operculum course=straight |operculum shape=conic;long-conic |perichaetial leaf shape=abruptly subulate;gradually acuminate |perigonial leaf shape=long-acuminate;short-acuminate;concave;ovate |peristome quantity=single |plant development=well-developed |propagule architecture=multicellular |propagule presence=absent |propagule reproduction=asexual |propagule texture=filamentous |rhizoid relief=papillose |row quantity=2;3 |segment fusion=free;united |segment quantity=2 |segment shape=filiform |seta architecture=twisted |seta coloration=yellow-orange;yellow;yellow-orange;yellow;purplish;red |seta orientation=erect |seta shape=elongate |spore relief=smooth;finely papillose |spore shape=globose |stem architecture=branched |stem atypical some measurement=3cm;4cm |stem some measurement=1cm;3cm |tooth quantity=16 |tuft density=dense |turf density=dense |wall arrangement=scattered |wall width=thin |whole_organism architecture or fragility=loose |whole_organism coloration=purple;tinged reddish-brown;green;dark green brownish green light green or yellow-green |whole_organism growth form=plant;plant |whole_organism reproduction=female;male }}<!-- -->[[Category:Treatment]][[Category:Ditrichaceae]] Templates used on this page: Template:Ditrichaceae (view source) Template:Treatment/AuthorLink (view source) Template:Treatment/Body (view source) Template:Treatment/Body/Maps (view source) Template:Treatment/ID (view source) Template:Treatment/Publication (view source) Template:Treatment/Reference (view source) Return to Ceratodon.