View source for Hedwigia ← Hedwigia You do not have permission to edit this page, for the following reason: The action you have requested is limited to users in the group: Users. You can view and copy the source of this page. {{Treatment/ID |accepted_name=Hedwigia |accepted_authority=P. Beauvois |publications={{Treatment/Publication |title=Mag. Encycl. |place=5: 304. 1804 |year=1804 }} |basionyms= |synonyms= |hierarchy=Hedwigiaceae;Hedwigia |hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Hedwigiaceae]]</div><div class="higher-taxon"><small>genus</small>[[Hedwigia]]</div></div> |etymology=For Johann Hedwig, 1730 – 1799, German bryologist and physician |volume=Volume 28 |mention_page=page 85, 88, 90, 642 |treatment_page=page 84 }}<!-- --><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism pubescence;whole_organism growth form"><b>Plants </b>large, sordid yellow-green, rarely bright green, pale buff to redbrown with age, sometimes hoary distally.</span> <span class="statement" id="st-d0_s1" data-properties="branch shape;branch presence"><b>Stems </b>with stoloniform-flagelliform branches absent.</span> <span class="statement" id="st-d0_s2" data-properties="leaf orientation;leaf arrangement;leaf arrangement;leaf architecture or arrangement or shape or vernation;leaf relief;leaf some measurement"><b>Leaves </b>erect to subsecund and imbricate, not undulate-plicate, occasionally somewhat striolate when dry, 1.5–3 mm;</span> <span class="statement" id="st-d0_s3" data-properties="margin orientation;margin shape;margin shape;margin orientation;margin orientation;margin orientation;margin orientation;margin architecture or shape">margins recurved or sometimes plane in proximal 1/2–2/3, plane, erect, or incurved in acumen, in larger leaves closely and irregularly dentate to spinulose-dentate in apex;</span> <span class="statement" id="st-d0_s4" data-properties="tip coloration">apex erect to patent when dry, widespreading and recurved or reflexed when moist, acute, acuminate, or subpiliferous, concolorous when muticous, or hyaline-white with pellucid tip, canaliculate-furrowed to tubular, papillose proximally, spinose-serrate medially, smooth apically;</span> <span class="statement" id="st-d0_s5" data-properties="apex orientation;apex orientation;apex orientation;apex orientation;apex shape;apex shape;apex coloration;apex coloration;apex architecture;apex shape;apex relief;apex architecture or shape;apex architecture or pubescence or relief;cell shape;cell shape;cell shape;cell shape;cell shape;cell relief;region width;region architecture;region coloration;row quantity">alar cells oblate, quadrate, or short-rectangular, smooth to simple-papillose, walls even to thick and porose, region concolorous or color similar to mid basal region, in several rows along margins;</span> <span class="statement" id="st-d0_s6" data-properties="mid basal laminal cell shape;papilla quantity;row quantity;wall width;wall architecture;region coloration;leaf insertion">mid basal laminal cells long-rectangular, papillae 4–7, in 1 row, walls strongly thick, moderately to strongly porose, region strongly yellow to red-orange across insertion or to 1/4–1/3 (–1/2) leaf length;</span> <span class="statement" id="st-d0_s7" data-properties="papilla architecture;papilla architecture;wall width;wall shape">medial cells 1- or multipapillose on both surfaces, papillae simple or multifid, walls thick, moderately to coarsely and irregularly porose-sinuate;</span> <span class="statement" id="st-d0_s8" data-properties="distal cell variability;distal cell relief">distal cells chlorophyllose and similar to medial cells, or echlorophyllose and differentiated, prorulose at distal cell ends, papillose;</span> <span class="statement" id="st-d0_s9" data-properties="">apical cells pellucid, less papillose.</span> <span class="statement" id="st-d0_s10" data-properties="apical cell coloration;apical cell relief;apical cell reproduction;apical cell reproduction"><b>Sexual </b>condition autoicous;</span> <span class="statement" id="st-d0_s11" data-properties="margin architecture;margin architecture">perichaetial leaves to 4 mm, margins entire or ciliate distally.</span> <span class="statement" id="st-d0_s12" data-properties="vaginulum architecture or pubescence or relief;vaginulum relief;paraphyse quantity;calyptra development"><b>Vaginula </b>with paraphyses few-to-many, smooth, occasionally papillose, often extending onto developing calyptra.</span> <span class="statement" id="st-d0_s13" data-properties="seta coloration;seta some measurement;seta fragility or size;seta prominence"><b>Seta </b>reddish-brown, 0.5–0.9 mm, stout, hidden by perichaetial leaves.</span> <span class="statement" id="st-d0_s14" data-properties="capsule prominence;capsule coloration;capsule reflectance;capsule coloration;capsule shape;capsule shape;capsule shape;capsule shape;capsule shape;capsule shape;capsule some measurement;capsule relief;capsule architecture;capsule architecture or pubescence or relief;mouth width"><b>Capsule </b>deeply immersed, brown proximally, shiny redbrown at mouth, turbinate-urceolate when dry, subglobose, short-ovoid, or obovoid when moist, 1 mm, longitudinally wrinkled to sharply sulcate when dry, smooth except for neck, mouth wide;</span> <span class="statement" id="st-d0_s15" data-properties="">stomata cryptoporic;</span> <span class="statement" id="st-d0_s16" data-properties="operculum shape;operculum architecture or shape">operculum planoconvex, sometimes subumbonate-apiculate.</span> <span class="statement" id="st-d0_s17" data-properties="calyptra shape;calyptra some measurement;calyptra pubescence;calyptra architecture"><b>Calyptra </b>conic-mitrate, 0.5–0.9 mm, covering apex of capsule, pilose or naked.</span> <span class="statement" id="st-d0_s18" data-properties="spore some measurement;spore relief"><b>Spores </b>19–30 µm, finely to somewhat coarsely vermiculate-papillose.</span><!-- -->{{Treatment/Body |distribution=Nearly worldwide |discussion=<p>Species 6 (3 in the flora).</p><!-- --><p>Hedwigia in North America was until recently considered as a single species, with numerous varieties and forms (G. N. Jones 1933). Although a hair-point is often ascribed to species of Hedwigia, the acumen may be elongated but there is never a hairlike process as in Pseudobraunia. Long, multicellular cilia on leaf margins are rare among bryophytes (B. Goffinet et al. 2008), but are present on the perichaetial leaves in H. ciliata and H. stellata.</p><!-- --><p>The type of Hedwigia integrifolia P. Beauvois was collected in North America. The name was used for a widespread species now considered Braunia imberbis (Smith) N. Dalton & D. G. Long. The type of H. integrifolia was examined by N. J. Dalton et al. (2012) and found unrelated to B. imberbis. Hedwigia integrifolia was synonymized with Hedwigidium imberbe (Smith) Bruch & Schimper by W. J. Hooker and T. Taylor (1818). Presently the application of the name H. integrifolia is awaiting further study, although G. N. Jones (1933) considered H. integrifolia a synonym of H. ciliata (B. H. Allen 2010).</p> |tables= |references= }}<!-- --><div class="treatment-key"> ==Key== <div class="treatment-key-group"> {| class="wikitable fna-keytable" |-id=key-0-1 |1 |Medial and distal laminal cells with papillae (1-)2-4 per cell, simple, sessile, or low-stalked, variously branched; perichaetial leaf margins ciliate distally; apical laminal cell obtuse to truncate, multipapillose-coronate, short-rhomboidal, (40-)50-75(-80) µm. |[[Hedwigia ciliata|Hedwigia ciliata]] |-id=key-0-1 |1 |Medial and distal laminal cells with papillae usually 1 per cell, stalked, strongly, irregularly branched; perichaetial leaf margins ciliate or entire distally; apical laminal cell sharply pointed, smooth or papillose, long-linear, (80-)120-175(-200) µm |[[#key-0-2| > 2]] |-id=key-0-2 |2 |Leaf apices wide-spreading to squarrose when dry; laminal cells in hyaline area with some papillae large, spinose, among shorter papillae; papillae on abaxial leaf surface with branches of unequal lengths; perichaetial leaf margins long-ciliate distally. |[[Hedwigia stellata|Hedwigia stellata]] |-id=key-0-2 |2 |Leaf apices erect when dry; laminal cells in hyaline area with papillae low, simple, in lines; papillae on abaxial leaf surface with branches usually of equal lengths; perichaetial leaf margins entire. |[[Hedwigia detonsa|Hedwigia detonsa]] |} </div></div><!-- -->{{#Taxon: name=Hedwigia |authority=P. Beauvois |rank=genus |parent rank=family |synonyms= |basionyms= |family=Hedwigiaceae |distribution=Nearly worldwide |reference=None |publication title=Mag. Encycl. |publication year=1804 |special status= |source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_132.xml |genus=Hedwigia |apex architecture=canaliculate-furrowed |apex architecture or pubescence or relief=smooth |apex architecture or shape=spinose-serrate |apex coloration=hyaline-white;concolorous |apex orientation=reflexed;recurved;widespreading;erect;patent |apex relief=papillose |apex shape=tubular;acuminate;acute |apical cell coloration=pellucid |apical cell relief=papillose |apical cell reproduction=autoicous;sexual |branch presence=absent |branch shape=stoloniform-flagelliform |calyptra architecture=naked |calyptra development=developing |calyptra pubescence=pilose |calyptra shape=conic-mitrate |calyptra some measurement=0.5mm;0.9mm |capsule architecture=sulcate |capsule architecture or pubescence or relief=smooth |capsule coloration=redbrown;brown |capsule prominence=immersed |capsule reflectance=shiny |capsule relief=wrinkled |capsule shape=obovoid;short-ovoid;obovoid;short-ovoid;subglobose;turbinate-urceolate |capsule some measurement=1 |cell relief=smooth;simple-papillose |cell shape=short-rectangular;quadrate;short-rectangular;quadrate;oblate |distal cell relief=papillose |distal cell variability=differentiated |leaf architecture or arrangement or shape or vernation=not undulate-plicate |leaf arrangement=imbricate;subsecund |leaf insertion=length |leaf orientation=erect |leaf relief=striolate |leaf some measurement=1.5mm;3mm |margin architecture=ciliate;entire |margin architecture or shape=irregularly dentate;spinulose-dentate |margin orientation=incurved;erect;incurved;erect;recurved |margin shape=plane;plane |mid basal laminal cell shape=long-rectangular |mouth width=wide |operculum architecture or shape=subumbonate-apiculate |operculum shape=planoconvex |papilla architecture=multifid;simple |papilla quantity=4;7 |paraphyse quantity=few;many |region architecture=porose |region coloration=strongly yellow;red-orange |region width=thick |row quantity=1;several |seta coloration=reddish-brown |seta fragility or size=stout |seta prominence=hidden |seta some measurement=0.5mm;0.9mm |spore relief=vermiculate-papillose |spore some measurement=19um;30um |tip coloration=pellucid |vaginulum architecture or pubescence or relief=smooth |vaginulum relief=papillose |wall architecture=porose |wall shape=porose-sinuate |wall width=thick;thick |whole_organism coloration=pale buff;redbrown |whole_organism growth form=plant |whole_organism pubescence=hoary |whole_organism size=large }}<!-- -->[[Category:Treatment]][[Category:Hedwigiaceae]] Templates used on this page: Template:Hedwigiaceae (view source) Template:Treatment/AuthorLink (view source) Template:Treatment/Body (view source) Template:Treatment/Body/Maps (view source) Template:Treatment/ID (view source) Template:Treatment/Publication (view source) Return to Hedwigia.