View source for Hilpertia

{{Treatment/ID
|accepted_name=Hilpertia
|accepted_authority=R. H. Zander
|publications={{Treatment/Publication
|title=Phytologia
|place=65: 427. 1989 ,
}}
|basionyms=
|synonyms=
|hierarchy=Pottiaceae;Pottiaceae subfam. Pottioideae;Hilpertia
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Pottiaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Pottiaceae subfam. Pottioideae]]</div><div class="higher-taxon"><small>genus</small>[[Hilpertia]]</div></div>
|etymology=For Friedrich Hilpert, b. 1907, German bryologist
|volume=Volume 27
|mention_page=page 265, 480, 608, 632
|treatment_page=page 631
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism coloration;whole_organism coloration;whole_organism growth form;cushion architecture or fragility"><b>Plants </b>in loose cushions, greenish brown distally, light-brown proximally.</span> <span class="statement" id="st-d0_s1" data-properties="stem some measurement;hyalodermi presence;scleroderm variability;central strand fusion"><b>Stems </b>to 1 cm, hyalodermis absent, sclerodermis not or weakly differentiated, central strand distinct;</span> <span class="statement" id="st-d0_s2" data-properties="axillary hair quantity;cell length or size;cell coloration;proximal cell coloration">axillary hairs to 8 cells long, all hyaline or the proximal one yellowbrown.</span> <span class="statement" id="st-d0_s3" data-properties=""><b>Stem-</b>leaves crowded, larger distally, appressed and tightly spiraled when dry, weakly spreading when moist, 1–1.4 (plus 0.3–0.6 of awn) mm;</span> <span class="statement" id="st-d0_s4" data-properties="leaf-stem arrangement;leaf-stem size;leaf-stem fixation or orientation;leaf-stem arrangement;leaf-stem orientation;leaf-stem distance;leaf-stem shape;adaxial surface shape;adaxial surface shape;base variability">ovate to circular, adaxial surface flat to quite concave, base not differentiated;</span> <span class="statement" id="st-d0_s5" data-properties="distal margin shape or vernation;distal margin architecture or shape;distal margin architecture or shape;cell size;margin shape or vernation">distal margins strongly revolute (to 2 ×), entire or broadly dentate above, cells of revolute margin enlarged, strongly chlorophyllose;</span> <span class="statement" id="st-d0_s6" data-properties="apex shape;apex coloration">apex broadly acute, hyaline in an apical triangle;</span> <span class="statement" id="st-d0_s7" data-properties="costa size or width;costa width;costa architecture;awn coloration;lamina position;adaxial outgrowth quantity;adaxial superficial cell shape;row quantity">costa narrow but broader above, excurrent as a hyaline awn, lamina inserted laterally, adaxial outgrowths none, superficial adaxial cells long-rectangular, in 2–4 rows;</span> <span class="statement" id="st-d0_s8" data-properties="transverse-section shape;adaxial epidermis presence;band presence;guide cell quantity;layer quantity;hydroid strand presence;band presence;band shape;abaxial epidermis presence">transverse-section rounded, adaxial epidermis present, adaxial stereid band absent, guide cells 2 in 1 layer, hydroid strand present, abaxial stereid band present, rounded in cross-sectional shape, abaxial epidermis absent;</span> <span class="statement" id="st-d0_s9" data-properties="basal-cell variability;basal-cell shape;basal-cell architecture">basal-cells weakly differentiated, rectangular, thin-walled;</span> <span class="statement" id="st-d0_s10" data-properties="distal medial cell shape;distal medial cell shape;distal medial cell size;distal medial cell width;distal medial cell width;distal medial cell quantity;distal medial cell quantity">distal medial cells hexagonal to shortrectangular or rhomboid, rather large, (14–) 20–25 µm wide, 1–4: 1;</span> <span class="statement" id="st-d0_s11" data-properties="distal medial cell shape;distal medial cell relief;papilla shape;papilla size;papilla width;papilla atypical some measurement;papilla some measurement;papilla shape;papilla size;papilla width;papilla quantity;papilla shape;papilla size;papilla width;papilla presence;margin shape or vernation;cell-wall width;cell-wall architecture;superficial abaxial cell-wall width;superficial abaxial cell-wall shape">papillae absent medially, hollow-papillose on revolute margins, cell-walls thin to thickened and porose, superficial abaxial cell-walls thick, weakly convex superficially on both sides, cells of leaf apex rhomboid to fusiform, smooth.</span> <span class="statement" id="st-d0_s12" data-properties="cell shape;cell architecture or pubescence or relief;cell development;brood reproduction;proximal rhizoid coloration;proximal rhizoid shape"><b>Specialized </b>asexual reproduction by brood bodies borne on proximal rhizoids, brown, spheric to elliptic.</span> <span class="statement" id="st-d0_s13" data-properties="whole-organism reproduction;whole-organism reproduction;whole-organism reproduction;whole-organism reproduction;whole-organism reproduction;perigonium position or structure subtype;plant arrangement"><b>Sexual </b>condition polyoicous: synoicous, paroicous, autoicous or perigonia terminal on separate plants.</span> <span class="statement" id="st-d0_s14" data-properties="perichaetium position or structure subtype;interior leaf architecture or shape;interior leaf variability;interior leaf shape;margin variability;medial laminal cell shape;medial laminal cell architecture"><b>Perichaetia </b>terminal, interior leaves sheathing, usually differentiated, long-oval, margins usually somewhat differentiated, medial laminal cells rectangular, very thin-walled.</span> <span class="statement" id="st-d0_s15" data-properties="seta some measurement"><b>Seta </b>0.35–0.4 cm.</span> <span class="statement" id="st-d0_s16" data-properties="capsule dehiscence;theca arrangement or shape;theca shape;theca some measurement;row quantity;cell size;cell shape;cell architecture"><b>Capsule </b>stegocarpous, theca elliptic, occasionally weakly ventricose, 1.2–1.5 mm, annulus of 3 rows of smaller, quadrate, highly vesiculose cells;</span> <span class="statement" id="st-d0_s17" data-properties="operculum shape;peristome tooth quantity;peristome tooth length or size;peristome tooth arrangement or course or shape;peristome tooth architecture;peristome tooth quantity">operculum broadly short-conic to long-conic, peristome teeth 32, long, linear, twisted counterclockwise about 1/2 turn.</span> <span class="statement" id="st-d0_s18" data-properties="calyptra shape"><b>Calyptra </b>cucullate.</span> <span class="statement" id="st-d0_s19" data-properties="spore some measurement;spore coloration;spore coloration"><b>Spores </b>13–16 µm. KOH laminal color reaction red.</span><!--

-->{{Treatment/Body
|distribution=Canada;Europe;Asia
|discussion=<p>Species 1.</p><!--
--><p>Hilpertia is unique among the Pottioideae in the elaboration of a photosynthetic organ consisting of laminal marginal cells rolled in a spiral tube, thin-walled and hollow-papillose. The species is characteristic of cold steppe habitats.</p>
|tables=
|references=
}}<!--

--><!--

-->{{#Taxon:
name=Hilpertia
|author=Richard H. Zander
|authority=R. H. Zander
|rank=genus
|parent rank=subfamily
|synonyms=
|basionyms=
|family=Pottiaceae
|distribution=Canada;Europe;Asia
|reference=None
|publication title=Phytologia
|publication year=
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27_930.xml
|subfamily=Pottiaceae subfam. Pottioideae
|genus=Hilpertia
|abaxial epidermis presence=absent
|adaxial epidermis presence=absent
|adaxial outgrowth quantity=0
|adaxial superficial cell shape=long-rectangular
|adaxial surface shape=concave;flat to quite
|apex coloration=hyaline
|apex shape=acute
|awn coloration=hyaline
|axillary hair quantity=0;8
|band presence=absent;absent
|band shape=rounded
|basal-cell architecture=thin-walled
|basal-cell shape=rectangular
|basal-cell variability=differentiated
|base variability=not differentiated
|brood reproduction=asexual
|calyptra shape=cucullate
|capsule dehiscence=stegocarpous
|cell architecture=vesiculose
|cell architecture or pubescence or relief=smooth
|cell coloration=hyaline
|cell development=specialized
|cell length or size=long
|cell shape=quadrate;rhomboid;fusiform
|cell size=smaller;enlarged
|cell-wall architecture=porose
|cell-wall width=thin;thickened
|central strand fusion=distinct
|costa architecture=excurrent
|costa size or width=narrow
|costa width=broader
|cushion architecture or fragility=loose
|distal margin architecture or shape=dentate;entire
|distal margin shape or vernation=revolute
|distal medial cell quantity=1;1;4
|distal medial cell relief=hollow-papillose
|distal medial cell shape=hexagonal;rhomboid;hexagonal
|distal medial cell size=large
|distal medial cell width=20um;25um
|guide cell quantity=2
|hyalodermi presence=absent
|hydroid strand presence=absent
|interior leaf architecture or shape=sheathing
|interior leaf shape=long-oval
|interior leaf variability=differentiated
|lamina position=inserted
|layer quantity=1
|leaf-stem arrangement=spiraled;crowded
|leaf-stem distance=1mm;1.4mm
|leaf-stem fixation or orientation=appressed
|leaf-stem orientation=spreading
|leaf-stem shape=ovate;circular
|leaf-stem size=larger
|margin shape or vernation=revolute;revolute
|margin variability=differentiated
|medial laminal cell architecture=thin-walled
|medial laminal cell shape=rectangular
|operculum shape=broadly short-conic;long-conic
|papilla atypical some measurement=14um;20um
|papilla presence=absent
|papilla quantity=1;4
|papilla shape=rhomboid;rhomboid;rhomboid
|papilla size=large;large;large
|papilla some measurement=20um;25um
|papilla width=wide;wide;wide
|perichaetium position or structure subtype=terminal
|perigonium position or structure subtype=terminal
|peristome tooth architecture=twisted
|peristome tooth arrangement or course or shape=linear
|peristome tooth length or size=long
|peristome tooth quantity=1/2;32
|plant arrangement=separate
|proximal cell coloration=yellowbrown
|proximal rhizoid coloration=brown
|proximal rhizoid shape=spheric;elliptic
|row quantity=3;2;4
|scleroderm variability=differentiated
|seta some measurement=0.35cm;0.4cm
|spore coloration=red;laminal
|spore some measurement=13um;16um
|stem some measurement=0cm;1cm
|superficial abaxial cell-wall shape=convex
|superficial abaxial cell-wall width=thick
|theca arrangement or shape=elliptic
|theca shape=ventricose
|theca some measurement=1.2mm;1.5mm
|transverse-section shape=rounded
|whole-organism reproduction=autoicous;paroicous;synoicous;polyoicous;sexual
|whole_organism coloration=light-brown;greenish brown
|whole_organism growth form=plant
}}<!--

-->[[Category:Treatment]][[Category:Pottiaceae subfam. Pottioideae]]