View source for Imbribryum

{{Treatment/ID
|accepted_name=Imbribryum
|accepted_authority=N. Pedersen
|publications={{Treatment/Publication
|title=Bryologist
|place=108: 449. 2005
|year=2005
}}
|basionyms=
|synonyms=
|hierarchy=Bryaceae;Imbribryum
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Bryaceae]]</div><div class="higher-taxon"><small>genus</small>[[Imbribryum]]</div></div>
|etymology=Latin imbrex, roof tile, and Greek bryon, moss, alluding to strongly overlapping leaves
|volume=Volume 28
|mention_page=page 11, 119, 143, 144, 658, 659
|treatment_page=page 142
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism growth form;turf architecture;turf density"><b>Plants </b>small to medium-sized, in open to dense turfs, green, yellow-green, red-green, or red.</span> <span class="statement" id="st-d0_s1" data-properties="stem atypical some measurement;stem some measurement;stem architecture;stem architecture or shape;stem architecture or shape;stem architecture"><b>Stems </b>0.5–3 (–5) cm, evenly foliate, julaceous or not, often branching;</span> <span class="statement" id="st-d0_s2" data-properties="rhizoid quantity;rhizoid presence;rhizoid position">rhizoids few, micronemata present at base of stem, occasionally in clusters along stem, macronemata axillary.</span> <span class="statement" id="st-d0_s3" data-properties="leaf arrangement;leaf orientation;leaf shape;leaf shape;leaf some measurement"><b>Leaves </b>strongly imbricate when dry, erect when moist, ovate to lanceolate or rarely lingulate or triangular, flat or concave, 1–3 mm;</span> <span class="statement" id="st-d0_s4" data-properties="base shape">base rarely decurrent;</span> <span class="statement" id="st-d0_s5" data-properties="margin shape;limbidium shape or architecture;limbidium architecture;limbidium presence;limbidium presence">margins plane to strongly revolute, entire to serrulate distally, 1-stratose, limbidium absent or occasionally present;</span> <span class="statement" id="st-d0_s6" data-properties="apex shape">apex obtuse to acuminate;</span> <span class="statement" id="st-d0_s7" data-properties="costa architecture;awn presence;layer quantity">costa percurrent to short-excurrent with awn present, guide cells usually in 1 layer;</span> <span class="statement" id="st-d0_s8" data-properties="cell variability">alar cells not differentiated;</span> <span class="statement" id="st-d0_s9" data-properties="laminal areolation variability">laminal areolation heterogeneous;</span> <span class="statement" id="st-d0_s10" data-properties="proximal laminal cell shape;proximal laminal cell shape;proximal laminal cell height or length or size;proximal laminal cell atypical quantity;proximal laminal cell quantity;proximal laminal cell quantity">proximal laminal cells abruptly quadrate or rectangular, shorter than more distal cells, 1–2 (–3):1;</span> <span class="statement" id="st-d0_s11" data-properties="medial and distal cell shape;medial and distal cell shape;medial and distal cell atypical quantity;medial and distal cell quantity;medial and distal cell quantity">medial and distal cells vermicular to rhomboidal or hexagonal, usually 3–6 (–8):1, walls thin to incrassate, rarely porose.</span> <span class="statement" id="st-d0_s12" data-properties="rhizoidal reproduction;rhizoidal presence;tuber presence"><b>Specialized </b>asexual reproduction by rhizoidal tubers or absent.</span> <span class="statement" id="st-d0_s13" data-properties="wall width;wall size;wall architecture;wall development;wall reproduction;wall reproduction"><b>Sexual </b>condition dioicous;</span> <span class="statement" id="st-d0_s14" data-properties="perigonium position or structure subtype;perichaetium position or structure subtype">perigonia and perichaetia terminal;</span> <span class="statement" id="st-d0_s15" data-properties="perigonial and perichaetial leaf variability;outer perichaetial leaf size;inner leaf size">perigonial and perichaetial leaves not much differentiated, outer perichaetial leaves larger, inner leaves smaller than leaves of innovations.</span> <span class="statement" id="st-d0_s16" data-properties="seta quantity;seta course;seta course"><b>Seta </b>usually single, ± straight or somewhat flexuose.</span> <span class="statement" id="st-d0_s17" data-properties="capsule orientation;capsule orientation;capsule shape;capsule some measurement"><b>Capsule </b>inclined or nutant, pyriform to clavate, 2–6 mm;</span> <span class="statement" id="st-d0_s18" data-properties="hypophysis size;hypophysis shape;hypophysis relief">hypophysis slender, not inflated or rugose;</span> <span class="statement" id="st-d0_s19" data-properties="operculum shape;operculum architecture or shape">operculum convex, apiculate;</span> <span class="statement" id="st-d0_s20" data-properties="">peristome double;</span> <span class="statement" id="st-d0_s21" data-properties="exostome coloration;tooth development;tooth shape">exostome yellowish, teeth well developed, lanceolate;</span> <span class="statement" id="st-d0_s22" data-properties="endostome fusion;basal membrane height;segment development;perforation shape;cilium length or size;cilium architecture;cilium shape">endostome not adherent to exostome, basal membrane high, segments well developed, with ovate perforations, cilia long, appendiculate or sometimes nodose.</span> <span class="statement" id="st-d0_s23" data-properties="spore arrangement;spore atypical some measurement;spore some measurement;spore relief;spore coloration;spore coloration"><b>Spores </b>shed singly, 10–18 (–20) µm, smooth to papillose, yellowish or yellowbrown.</span><!--

-->{{Treatment/Body
|distribution=Nearly worldwide;except Antarctica;tropical;subtropical;temperate to boreal regions;rare in arctic-alpine climates
|discussion=<p>Species ca. 40 (6 in the flora).</p><!--
--><p>Species of Imbribryum are relatively large, with elongate, evenly foliate stems and strongly imbricate leaves. The plants commonly grow on soil or rock, often associated with waterfalls, seeps and springs, or splashed rock. The plants have stout setae 1–3 cm in length; the capsules have well-developed peristomes. The genus is especially well represented in the mountains of New Guinea, the Neotropics, southeast Asia, and Africa. Most species produce spheric rhizoidal tubers; the presence of leaf axil bulbils, reported from European material of I. gemmiparum, has not been confirmed for North American collections. Many species are difficult to identify as they are morphologically variable, especially I. alpinum and I. muehlenbeckii. There is one undescribed species in the western United States, discussed under I. alpinum.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Leaves strongly imbricate when dry; distal laminal cells sometimes vermicular, 6-8:1; stems not or somewhat julaceous
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Leaves strongly to somewhat loosely imbricate; distal laminal cells not vermicular, usually 3-6:1; stems sometimes julaceous
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Costae percurrent to short-excurrent, awns stout; limbidium absent; proximal laminal cells quadrate to short-rectangular; capsule necks short.
|[[Imbribryum alpinum|Imbribryum alpinum]]
|-id=key-0-2
|2
|Costae short- to moderately long-excurrent, awns slender; limbidium present; proximal laminal cells rectangular; capsule necks long.
|[[Imbribryum microchaeton|Imbribryum microchaeton]]
|-id=key-0-3
|3
|Stems strongly julaceous; leaves purple-red to brown-red, rarely dull olive green, strongly concave; apices rounded-obtuse to broadly acute, cucullate; costae percurrent; distal laminal cell walls strongly incrassate, oblique to costa.
|[[Imbribryum miniatum|Imbribryum miniatum]]
|-id=key-0-3
|3
|Stems not or weakly julaceous (except I. muehlenbeckii); leaves red, green, or yellow, concave to flat; apices rounded, acute, or acuminate, not cucullate; costae not reaching apex to excurrent; distal laminal cell walls thin to firm, parallel to costa
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Leaves weakly concave or flat, ovate-lanceolate; apices acute to acuminate; costae short-excurrent, awns slender, sometimes denticulate.
|[[Imbribryum mildeanum|Imbribryum mildeanum]]
|-id=key-0-4
|4
|Leaves distinctly concave, ovate; apices rounded to acute; costae not reaching apex or rarely percurrent, awns absent
|[[#key-0-5| > 5]]
|-id=key-0-5
|5
|Leaves green, sometimes yellowish, soft, loosely set, somewhat distant proximally along stem; proximal laminal cells short-rectangular, occasionally with scattered quadrate cells.
|[[Imbribryum gemmiparum|Imbribryum gemmiparum]]
|-id=key-0-5
|5
|Leaves red or red-green, sometimes dark green, rigid, imbricate, crowded; proximal laminal cells quadrate, with scattered short-rectangular cells.
|[[Imbribryum muehlenbeckii|Imbribryum muehlenbeckii]]
|}
</div></div><!--

-->{{#Taxon:
name=Imbribryum
|author=John R. Spence
|authority=N. Pedersen
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Bryaceae
|distribution=Nearly worldwide;except Antarctica;tropical;subtropical;temperate to boreal regions;rare in arctic-alpine climates
|reference=None
|publication title=Bryologist
|publication year=2005
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_228.xml
|genus=Imbribryum
|apex shape=obtuse;acuminate
|awn presence=absent
|basal membrane height=high
|base shape=decurrent
|capsule orientation=nutant;inclined
|capsule shape=pyriform;clavate
|capsule some measurement=2mm;6mm
|cell variability=not differentiated
|cilium architecture=appendiculate
|cilium length or size=long
|cilium shape=nodose
|costa architecture=percurrent;short-excurrent
|endostome fusion=not adherent
|exostome coloration=yellowish
|hypophysis relief=rugose
|hypophysis shape=not inflated
|hypophysis size=slender
|inner leaf size=smaller
|laminal areolation variability=heterogeneous
|layer quantity=1
|leaf arrangement=imbricate
|leaf orientation=erect
|leaf shape=ovate;lanceolate or rarely lingulate or triangular flat or concave
|leaf some measurement=1mm;3mm
|limbidium architecture=1-stratose
|limbidium presence=absent;absent
|limbidium shape or architecture=entire to serrulate
|margin shape=plane;strongly revolute
|medial and distal cell atypical quantity=6;8
|medial and distal cell quantity=1;3;6
|medial and distal cell shape=hexagonal;rhomboidal
|operculum architecture or shape=apiculate
|operculum shape=convex
|outer perichaetial leaf size=larger
|perforation shape=ovate
|perichaetium position or structure subtype=terminal
|perigonial and perichaetial leaf variability=differentiated
|perigonium position or structure subtype=terminal
|proximal laminal cell atypical quantity=2;3
|proximal laminal cell height or length or size=shorter
|proximal laminal cell quantity=1;1;2
|proximal laminal cell shape=rectangular;quadrate
|rhizoid position=axillary
|rhizoid presence=absent
|rhizoid quantity=few
|rhizoidal presence=absent
|rhizoidal reproduction=asexual
|segment development=developed
|seta course=flexuose;straight
|seta quantity=single
|spore arrangement=singly
|spore atypical some measurement=18um;20um
|spore coloration=yellowbrown;yellowish
|spore relief=smooth;papillose
|spore some measurement=10um;18um
|stem architecture=branching;foliate
|stem architecture or shape=not;julaceous
|stem atypical some measurement=3cm;5cm
|stem some measurement=0.5cm;3cm
|tooth development=developed
|tooth shape=lanceolate
|tuber presence=absent
|turf architecture=open
|turf density=dense
|wall architecture=porose
|wall development=specialized
|wall reproduction=dioicous;sexual
|wall size=incrassate
|wall width=thin
|whole_organism coloration=red;red-green;red;red-green;yellow-green;green
|whole_organism growth form=plant
|whole_organism size=small;medium-sized
}}<!--

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