View source for Meiotrichum lyallii

{{Treatment/ID
|accepted_name=Meiotrichum lyallii
|accepted_authority=(Mitten) G. L. Smith
|publications={{Treatment/Publication
|title=Bryologist
|place=95: 271. 1992,
}}
|basionyms={{Treatment/ID/Basionym
|name=Polytrichadelphus lyallii
|authority=Mitten
|rank=species
|publication_title=J. Linn. Soc., Bot.
|publication_place=8: 49, plate 8. 1864
}}
|synonyms={{Treatment/ID/Synonym
|name=Polytrichastrum lyallii
|authority=(Mitten) G. L. Smith
|rank=species
}} {{Treatment/ID/Synonym
|name=Polytrichum angustidens
|authority=H. Lindberg
|rank=species
}} {{Treatment/ID/Synonym
|name=Polytrichum lyallii
|authority=(Mitten) Kindberg
|rank=species
}}
|hierarchy=Polytrichaceae;Meiotrichum;Meiotrichum lyallii
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Polytrichaceae]]</div><div class="higher-taxon"><small>genus</small>[[Meiotrichum]]</div><div class="higher-taxon"><small>species</small>[[Meiotrichum lyallii]]</div></div>
|volume=Volume 27
|mention_page=page 126, 142
|treatment_page=page 141
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism fragility;whole_organism coloration;whole_organism growth form;tuft architecture or fragility"><b>Plants </b>robust, in loose tufts or growing among other mosses, dark green to reddish-brown when old.</span> <span class="statement" id="st-d0_s1" data-properties="stem height;stem height;stem orientation;stem architecture;stem architecture;stem architecture;stem architecture;stem architecture"><b>Stems </b>2–3 (–7) cm high, erect, bracteate proximally, abruptly leafy distally, simple or more typically fastigiately branched from near the base and sparingly branched distally.</span> <span class="statement" id="st-d0_s2" data-properties="leaf arrangement;leaf orientation;leaf shape;leaf shape;leaf orientation;leaf orientation;leaf atypical some measurement;leaf some measurement"><b>Leaves </b>densely imbricate, ± spreading and subtubular when dry, plane and broadly recurved when moist, not reflexed at the hinge, (4–) 7–10 mm;</span> <span class="statement" id="st-d0_s3" data-properties="sheath some measurement;sheath architecture or shape;sheath architecture;marginal strip size or width;hinge-bud shape;portion shape">sheath 1.5–2 mm, entire, hyaline-margined, with a narrow marginal strip of transversely elongate hinge-cells in the tapering portion;</span> <span class="statement" id="st-d0_s4" data-properties="spine height or length or size">limb ending in a short spine;</span> <span class="statement" id="st-d0_s5" data-properties="marginal lamina orientation;marginal lamina architecture;tooth cell quantity;tooth cell atypical quantity;tooth cell quantity">marginal lamina erect, 1-stratose, the marginal teeth consisting of a single larger tooth cell and 1 (–2) smaller basal-cells;</span> <span class="statement" id="st-d0_s6" data-properties="costa architecture or pubescence or relief;tip shape">costa smooth on the back or weakly toothed at the tip;</span> <span class="statement" id="st-d0_s7" data-properties="lamella quantity;lamella quantity;bud height;bud architecture or shape;wall fusion;wall size or width;cuticle coloration or pubescence or relief;cuticle relief;cuticle shape;cuticle shape;section size;lumen relief">lamellae 36–38, 5–6 cells high, entire in profile, the marginal cells with only the free wall thickened, the cuticle irregularly striate and pitted, in section somewhat larger, ovate, appearing finely papillose, the lumen trapezoidal;</span> <span class="statement" id="st-d0_s8" data-properties="sheath bud length;sheath bud width;sheath bud quantity;sheath bud quantity;bud some measurement;bud reproduction;back shape;whole_organism reproduction;whole_organism arrangement;whole_organism growth form;portion architecture;portion reproduction;inflorescence atypical quantity;inflorescence quantity;inflorescence reproduction;leaf architecture">sheath cells 46–66 × 12.5–16 µm, 4–6: 1, cells on the back of limb ± uniformly subquadrate, 11–18 µm. <b>Sexual</b> condition: male plants with a ± abrupt transition to a spicate fertile portion consisting of 2 (–7) male inflorescences in series, separated by bracteate leaves;</span> <span class="statement" id="st-d0_s9" data-properties="sheath bud length;sheath bud width;sheath bud quantity;perigonial bract shape;perigonial bract architecture or shape;perigonial bract architecture or shape;perigonial bract architecture;perigonial bract arrangement;cup shape">perigonial bracts ovate, apiculate, entire, hyaline-margined, broadly overlapping, forming a cylindrical cup;</span> <span class="statement" id="st-d0_s10" data-properties="sheath bud length;sheath bud width;sheath bud quantity;paraphyse shape;paraphyse length">paraphyses clavate, 2-seriate for more than 1/2 their length;</span> <span class="statement" id="st-d0_s11" data-properties="sheath bud length;sheath bud width;sheath bud quantity;perichaetial leaf orientation;perichaetial leaf architecture or shape">perichaetial leaves erect, long-sheathing, the bases surrounding the seta at base.</span> <span class="statement" id="st-d0_s12" data-properties="seta position relational;seta reproduction;seta reproduction"><b>Sexual </b>condition dioicous;</span> <span class="statement" id="st-d0_s13" data-properties="whole_organism reproduction;whole_organism arrangement;whole_organism growth form;portion shape;portion duration;portion architecture;portion reproduction;inflorescence atypical quantity;inflorescence quantity;inflorescence reproduction;leaf architecture">male plants with a ± abrupt transition to a spicate fertile portion consisting of 2 (–7) male inflorescences in series, separated by bracteate leaves;</span> <span class="statement" id="st-d0_s14" data-properties="perigonial bract shape;perigonial bract architecture or shape;perigonial bract architecture or shape;perigonial bract architecture;perigonial bract arrangement;cup shape">perigonial bracts ovate, apiculate, entire, hyaline-margined, broadly overlapping, forming a cylindrical cup;</span> <span class="statement" id="st-d0_s15" data-properties="paraphyse shape;paraphyse length">paraphyses clavate, 2-seriate for more than 1/2 their length;</span> <span class="statement" id="st-d0_s16" data-properties="perichaetial leaf orientation;seta position relational">perichaetial leaves erect, the bases surrounding the seta at base.</span> <span class="statement" id="st-d0_s17" data-properties="seta some measurement;seta fragility or size;seta coloration;seta quantity;sporophyte quantity"><b>Seta </b>3–6 cm, stout, yellowish, 1 per perichaetium, but often several sporophytes per plant, one to a branch.</span> <span class="statement" id="st-d0_s18" data-properties="capsule some measurement;capsule coloration;capsule orientation;capsule width;capsule shape;capsule shape;capsule quantity;uppermost angle prominence;lower angle prominence"><b>Capsule </b>3–4 mm, brownish to reddish-brown, inclined to horizontal, widest near the base and tapering to the mouth, often ± bilaterally compressed, the two uppermost angles often more prominent and forming a crest, the lower angles less pronounced;</span> <span class="statement" id="st-d0_s19" data-properties="exothecial bud pubescence or shape;exothecial bud length;exothecial bud width;apophysis fusion;superficial stoma quantity">exothecial cells moderately bulging, 10.5–13.5 × 6–7.5 µm; apophysis distinct, with 30–40 superficial stomata;</span> <span class="statement" id="st-d0_s20" data-properties="operculum shape;operculum some measurement;beak fragility or size;beak shape">operculum hemispherical, with a stout, hooked beak, about 1.5 mm;</span> <span class="statement" id="st-d0_s21" data-properties="peristome coloration;peristome shape;peristome quantity;tooth quantity;tooth size;tooth shape">peristome pale, divided to 0.3, the teeth 64, slender and tapering.</span> <span class="statement" id="st-d0_s22" data-properties="calyptra shape;calyptra pubescence;hair quantity"><b>Calyptra </b>cucullate, hispid at the tip, with few hairs.</span> <span class="statement" id="st-d0_s23" data-properties="spore some measurement;spore relief"><b>Spores </b>9–13 µm, finely papillose.</span><!--

-->{{Treatment/Body
|habitat=Banks, thin soil over boulders in open woods, also on moist shaded cliffs, boulders
|elevation=moderate to high elevations (900-3300 m)
|distribution=B.C.;Yukon;Alaska;Ariz.;Calif.;Colo.;Idaho;Mont.;Nev.;N.Mex.;Oreg.;Utah;Wash.;Wyo.
|discussion=<p>The habit of Meiotrichum lyallii is characteristic, the stems short and fastigiately branched from near the base, bearing several sporophytes, one to a branch. The pale, slender, tapering peristome teeth are also distinctive. The androecia can take the form of spicate “cones,” with several consecutive male inflorescences in succession, not reverting to normal vegetative leaves in the interval between inflorescences. When sporophytes are present, M. lyallii is easily recognized, but can be mistaken for Polytrichastrum alpinum when sterile. Both have serrate-toothed leaves with sloping shoulders and similar-appearing lamellae, but in M. lyallii the cells appear papillose in section but are actually finely striolate and pitted, whereas in P. alpinum the cells are papillose, with distinct rounded papillae.</p>
|tables=
|references=
}}<!--

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-->{{#Taxon:
name=Meiotrichum lyallii
|authority=(Mitten) G. L. Smith
|rank=species
|parent rank=genus
|synonyms=Polytrichastrum lyallii;Polytrichum angustidens;Polytrichum lyallii
|basionyms=Polytrichadelphus lyallii
|family=Polytrichaceae
|habitat=Banks, thin soil over boulders in open woods, also on moist shaded cliffs, boulders
|elevation=moderate to high elevations (900-3300 m)
|distribution=B.C.;Yukon;Alaska;Ariz.;Calif.;Colo.;Idaho;Mont.;Nev.;N.Mex.;Oreg.;Utah;Wash.;Wyo.
|reference=None
|publication title=Bryologist
|publication year=
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27_174.xml
|genus=Meiotrichum
|species=Meiotrichum lyallii
|apophysis fusion=distinct
|back shape=subquadrate
|beak fragility or size=stout
|beak shape=hooked
|bud architecture or shape=entire
|bud height=high
|bud reproduction=sexual
|bud some measurement=11um;18um
|calyptra pubescence=hispid
|calyptra shape=cucullate
|capsule coloration=brownish;reddish-brown
|capsule orientation=inclined;horizontal
|capsule quantity=2
|capsule shape=compressed;tapering
|capsule some measurement=3mm;4mm
|capsule width=widest
|costa architecture or pubescence or relief=smooth
|cup shape=cylindrical;cylindrical
|cuticle coloration or pubescence or relief=striate
|cuticle relief=pitted
|cuticle shape=trapezoidal;ovate
|exothecial bud length=10.5um;13.5um
|exothecial bud pubescence or shape=bulging
|exothecial bud width=6um;7.5um
|hair quantity=few
|hinge-bud shape=elongate
|inflorescence atypical quantity=2;7
|inflorescence quantity=2;2
|inflorescence reproduction=male;male
|lamella quantity=5;6
|leaf architecture=bracteate;bracteate
|leaf arrangement=imbricate
|leaf atypical some measurement=4mm;7mm
|leaf orientation=not reflexed;recurved;spreading
|leaf shape=plane;subtubular
|leaf some measurement=7mm;10mm
|lower angle prominence=pronounced
|lumen relief=papillose
|marginal lamina architecture=1-stratose
|marginal lamina orientation=erect
|marginal strip size or width=narrow
|operculum shape=hemispherical
|operculum some measurement=1.5
|paraphyse length=2-seriate;2-seriate
|paraphyse shape=clavate;clavate
|perichaetial leaf architecture or shape=long-sheathing
|perichaetial leaf orientation=erect;erect
|perigonial bract architecture=hyaline-margined;hyaline-margined
|perigonial bract architecture or shape=entire;apiculate;entire;apiculate
|perigonial bract arrangement=overlapping;overlapping
|perigonial bract shape=ovate;ovate
|peristome coloration=pale
|peristome quantity=0.3
|peristome shape=divided
|portion architecture=spicate;spicate
|portion duration=transition
|portion reproduction=fertile;fertile
|portion shape=abrupt;tapering
|section size=larger
|seta coloration=yellowish
|seta fragility or size=stout
|seta position relational=surrounding;surrounding
|seta quantity=1
|seta reproduction=dioicous;sexual
|seta some measurement=3cm;6cm
|sheath architecture=hyaline-margined
|sheath architecture or shape=entire
|sheath bud length=46um;66um
|sheath bud quantity=4;6
|sheath bud width=12.5um;16um
|sheath some measurement=1.5mm;2mm
|spine height or length or size=short
|spore relief=papillose
|spore some measurement=9um;13um
|sporophyte quantity=several
|stem architecture=branched;branched;simple;leafy;bracteate
|stem height=2cm;3cm
|stem orientation=erect
|superficial stoma quantity=30;40
|tip shape=toothed
|tooth cell atypical quantity=1;2
|tooth cell quantity=1;single
|tooth quantity=64
|tooth shape=tapering
|tooth size=slender
|tuft architecture or fragility=loose
|uppermost angle prominence=prominent
|wall fusion=free
|wall size or width=thickened
|whole_organism arrangement=separated;separated
|whole_organism coloration=dark green;reddish-brown
|whole_organism fragility=robust
|whole_organism growth form=plant;plant;plant
|whole_organism reproduction=male;male
}}<!--

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