View source for Nyssa

{{Treatment/ID
|accepted_name=Nyssa
|accepted_authority=Linnaeus
|publications={{Treatment/Publication
|title=Sp. Pl.
|place=2: 1058. 1753
|year=1753
}}
|common_names=Tupelo
|basionyms=
|synonyms=
|hierarchy=Nyssaceae;Nyssa
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Nyssaceae]]</div><div class="higher-taxon"><small>genus</small>[[Nyssa]]</div></div>
|etymology=Classical Greek name for a water nymph, alluding to habitat
|volume=Volume 12
|mention_page=page 438, 443, 458
|treatment_page=page 459
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism growth form;whole_organism growth form"><b>Shrubs </b>or trees;</span> <span class="statement" id="st-d0_s1" data-properties="bark coloration;bark pubescence or relief;bark shape">bark gray brown, rough, ridged;</span> <span class="statement" id="st-d0_s2" data-properties="diaphragm dehiscence or orientation">twigs with transverse diaphragms;</span> <span class="statement" id="st-d0_s3" data-properties="bud season;bud architecture or pubescence">winter buds scaly.</span> <span class="statement" id="st-d0_s4" data-properties="petiole shape;petiole architecture"><b>Leaves:</b> petiole terete or winged;</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;base shape">blade usually elliptic to oblanceolate or obovate, rarely ovate, base cuneate to rounded.</span> <span class="statement" id="st-d0_s6" data-properties="pedicel quantity;pedicel presence;pedicel presence;pedicel reproduction;pedicel architecture;pedicel presence"><b>Pedicels:</b> staminate present or absent, bisexual and pistillate absent [present].</span> <span class="statement" id="st-d0_s7" data-properties="rim position"><b>Flowers:</b> sepals forming a low rim;</span> <span class="statement" id="st-d0_s8" data-properties="petal coloration">petals greenish to greenish white;</span> <span class="statement" id="st-d0_s9" data-properties="style shape;style shape">style subulate or conic.</span> <span class="statement" id="st-d0_s10" data-properties="drupe coloration;drupe shape;remnant duration"><b>Drupes </b>usually blue-black (sometimes reddish purple in N. aquatica; yellow, orange, or red in N. ogeche), topped by persistent remnants of sepals;</span> <span class="statement" id="st-d0_s11" data-properties="mesocarp texture;mesocarp taste">mesocarp juicy, acidic.</span><!--

-->{{Treatment/Body
|distribution=c;e North America;Mexico;Central America (Costa Rica);Central America (Panama);e Asia
|discussion=<p>Species ca. 13 (5 in the flora).</p><!--
--><p>Plant sexuality in Nyssa is complicated and often difficult to determine. R. E. Burckhalter (1992) described all North American species as dioecious, without elaboration. The most widely distributed species in the flora area, N. sylvatica, has both staminate and morphologically bisexual flowers and appears to be androdioecious, but is functionally dioecious, as anthers of bisexual flowers do not dehisce (R. H. Eyde 1963; M. L. Cipollini and E. W. Stiles 1991). Similarly, the Chinese N. yunnanensis W. Q. Yin ex H. N. Qin & Phengklai appears to be androdioecious but is functionally dioecious; in this case pollen from the morphologically bisexual flowers is inaperturate and inviable, rendering the trees that bear these flowers functionally pistillate (Sun B.-L. et al. 2009). Nyssa ogeche appears to be polygamodioecious and other North American species appear to be monoecious or polygamomonoecious, but their reproductive biology has not been examined carefully.</p><!--
--><p>Measurements of leaves refer to fully expanded ones from mid shoot (earlier leaves typically are smaller).</p>
|tables=
|references={{Treatment/Reference
|id=burckhalter1992a
|text=Burckhalter, R. E. 1992. The genus Nyssa (Cornaceae) in North America: A revision. Sida 15: 323–342.
}}{{Treatment/Reference
|id=wen1993a
|text=Wen, J. and T. F. Stuessy. 1993. The phylogeny and biogeography of Nyssa (Cornaceae). Syst. Bot. 18: 68–79.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Drupes 22–28 mm; ovaries hairy; staminate pedicels absent.
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Petioles 30–60 mm; drupes punctate, black to blue or reddish purple.
|[[Nyssa aquatica|Nyssa aquatica]]
|-id=key-0-2
|2
|Petioles 6–20(–35) mm; drupes smooth, yellow, orange, or red.
|[[Nyssa ogeche|Nyssa ogeche]]
|-id=key-0-1
|1
|Drupes 7–14 mm; ovaries glabrous; staminate pedicels present.
|[[#key-0-3| > 3]]
|-id=key-0-3
|3
|Shrubs or trees, 2–5 m; Florida panhandle region.
|[[Nyssa ursina|Nyssa ursina]]
|-id=key-0-3
|3
|Trees, 5–30 m; widespread in c, e North America, including Florida.
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Leaf blades herbaceous, obovate to elliptic, apices acute to acuminate, margins entire proximally and 0–1(–3)-toothed distally; bark ± regularly longitudinally and transversely fissured; pistillate and bisexual inflorescences (2–)3–5(–8)-flowered.
|[[Nyssa sylvatica|Nyssa sylvatica]]
|-id=key-0-4
|4
|Leaf blades subcoriaceous, usually oblanceolate to narrowly elliptic, rarely ovate, apices obtuse to acute, margins usually entire, rarely coarsely dentate distally; bark irregularly fissured; pistillate and bisexual inflorescences 1–3-flowered.
|[[Nyssa biflora|Nyssa biflora]]
|}
</div></div><!--

-->{{#Taxon:
name=Nyssa
|author=Gordon C. Tucker;Tracy J. Park
|authority=Linnaeus
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Nyssaceae
|distribution=c;e North America;Mexico;Central America (Costa Rica);Central America (Panama);e Asia
|reference=burckhalter1992a;wen1993a
|publication title=Sp. Pl.
|publication year=1753
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V12/V12_137.xml
|genus=Nyssa
|bark coloration=gray brown
|bark pubescence or relief=rough
|bark shape=ridged
|base shape=cuneate;rounded
|blade shape=ovate;usually elliptic;oblanceolate or obovate
|bud architecture or pubescence=scaly
|bud season=winter
|diaphragm dehiscence or orientation=transverse
|drupe coloration=blue-black
|drupe shape=topped
|mesocarp taste=acidic
|mesocarp texture=juicy
|pedicel architecture=pistillate
|pedicel presence=absent;absent;absent
|pedicel quantity=staminate
|pedicel reproduction=bisexual
|petal coloration=greenish;greenish white
|petiole architecture=winged
|petiole shape=terete
|remnant duration=persistent
|rim position=low
|style shape=conic;subulate
|whole_organism growth form=tree;shrub
}}<!--

-->[[Category:Treatment]][[Category:Nyssaceae]]