View source for Rhytidiadelphus

{{Treatment/ID
|accepted_name=Rhytidiadelphus
|accepted_authority=(Limpricht) Warnstorf
|publications={{Treatment/Publication
|title=Krypt.-Fl. Brandenburg
|place=2: 917. 1906
|year=1906
}}
|basionyms={{Treatment/ID/Basionym
|name=Hylocomium subg. Rhytidiadelphus
|authority=Limpricht
|rank=subgenus
|publication_title=Laubm. Deutschl.
|publication_place=3: 590. 1901
}}
|synonyms={{Treatment/ID/Synonym
|name=Hylocomiadelphus
|authority=
|rank=genus
}}
|hierarchy=Hylocomiaceae;Rhytidiadelphus
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Hylocomiaceae]]</div><div class="higher-taxon"><small>genus</small>[[Rhytidiadelphus]]</div></div>
|etymology=Genus Rhytidium and Greek adelphos, brother, alluding to relationship
|volume=Volume 28
|mention_page=page 316, 326, 331, 338, 642, 644, 645
|treatment_page=page 330
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="stem orientation;stem width;stem architecture;stem architecture;stem architecture"><b>Stems </b>creeping to ascending or erect, 2–8 mm wide across leafy stem, monopodial, irregularly and remotely branched to subpinnate;</span> <span class="statement" id="st-d0_s1" data-properties="paraphyllium presence">paraphyllia absent.</span> <span class="statement" id="st-d0_s2" data-properties="leaf-stem orientation;leaf-stem orientation;leaf-stem arrangement;leaf-stem arrangement;leaf-stem arrangement;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem shape;leaf-stem architecture or arrangement or shape or vernation;leaf-stem relief;leaf-stem distance"><b>Stem-</b>leaves erect-spreading to squarrose, squarrose-recurved, or falcate-secund, crowded or not, broadly ovate, ovate-triangular, or ovatelanceolate, sometimes plicate proximally, sometimes rugose, 2.1–4.9 mm;</span> <span class="statement" id="st-d0_s3" data-properties="base shape">base not or slightly decurrent;</span> <span class="statement" id="st-d0_s4" data-properties="margin architecture or shape;margin architecture or shape">margins serrulate to nearly entire proximally, serrulate distally;</span> <span class="statement" id="st-d0_s5" data-properties="apex shape">apex acuminate;</span> <span class="statement" id="st-d0_s6" data-properties="">costa double, short or to 2/3 leaf length;</span> <span class="statement" id="st-d0_s7" data-properties="costa quantity;cell variability;cell height or length or size;cell width">alar cells not differentiated, or shorter, wider than basal-cells;</span> <span class="statement" id="st-d0_s8" data-properties="laminal cell architecture or pubescence or relief;laminal cell architecture or pubescence or relief">laminal cells smooth or coarsely prorate.</span> <span class="statement" id="st-d0_s9" data-properties="branch leaf shape"><b>Branch </b>leaves ovate to narrowly lanceolate.</span> <span class="statement" id="st-d0_s10" data-properties="capsule orientation"><b>Capsule </b>horizontal;</span> <span class="statement" id="st-d0_s11" data-properties="operculum shape">operculum conic;</span> <span class="statement" id="st-d0_s12" data-properties="exostome tooth pubescence;exostome tooth architecture or coloration or relief">exostome teeth cross-striolate to reticulate;</span> <span class="statement" id="st-d0_s13" data-properties="endostome segment architecture">endostome segments broadly perforate.</span><!--

-->{{Treatment/Body
|distribution=North America;Eurasia;Atlantic Islands;Pacific Islands (New Zealand);Australia (Tasmania);cool temperate and boreal zones
|discussion=<p>Species 5 (5 in the flora).</p><!--
--><p>In recent years, M. S. Ignatov and E. A. Ignatova (2003+, vol. 2) and R. Ochyra and A. Stebel (2008) have advocated placing Rhytidiadelphus triquetrus in its own monospecific genus. Although it is morphologically the most distinctive of the five species of Rhytidiadelphus, evolutionary relationships among the five are not clear. Some molecular studies have suggested a less than close relationship between R. squarrosus and R. triquetrus, but these studies were not directly focused on Hylocomiaceae. On the other hand, the analysis of morphological and anatomical evidence by L. Hedenäs (2004) in his cladistic study of Hylocomiaceae revealed that R. loreus and R. triquetrus share a more recent common ancestor than either does with the other three species. Likewise, ITS and ISSR molecular markers employed by A. Vanderpoorten et al. (2003) delineated two pairs of closely related species: R. loreus with R. triquetrus and R. squarrosus with R. subpinnatus. Until further study clarifies relationships within both Rhytidiadelphus and Hylocomiaceae, it seems best to maintain the traditional circumscription of the genus.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Leaves plicate; alar cells not or slightly differentiated
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Leaves not plicate; alar cells differentiated
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Costae 1/2 -2/3 leaf length; medial laminal cells coarsely prorate; stem leaf apices gradually narrowed, acumina broad, flat (occasionally rugose).
|[[Rhytidiadelphus triquetrus|Rhytidiadelphus triquetrus]]
|-id=key-0-2
|2
|Costae less than 1/4 leaf length; medial laminal cells smooth; stem leaf apices abruptly narrowed, acumina slender, channeled.
|[[Rhytidiadelphus loreus|Rhytidiadelphus loreus]]
|-id=key-0-3
|3
|Costae 1/3-1/2 leaf length; alar cell walls thick, porose; stem leaf acumina short.
|[[Rhytidiadelphus japonicus|Rhytidiadelphus japonicus]]
|-id=key-0-3
|3
|Costae to 1/3 leaf length; alar cell walls thin, not porose; stem leaf acumina long
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Stem leaves not crowded (stem visible between leaves) except at stem apices, erect-spreading to squarrose; bases not or scarcely sheathing; leaves broadly ovate-triangular; stems irregularly branched to pinnate; moist, usually shaded sites.
|[[Rhytidiadelphus subpinnatus|Rhytidiadelphus subpinnatus]]
|-id=key-0-4
|4
|Stem leaves crowded (stem concealed by leaves), strongly squarrose-recurved; bases subsheathing; leaves ovate; stems remotely branched to irregularly pinnate; well-drained, open to partially shaded sites, near coast or in grassy, disturbed habitats.
|[[Rhytidiadelphus squarrosus|Rhytidiadelphus squarrosus]]
|}
</div></div><!--

-->{{#Taxon:
name=Rhytidiadelphus
|authority=(Limpricht) Warnstorf
|rank=genus
|parent rank=family
|synonyms=Hylocomiadelphus
|basionyms=Hylocomium subg. Rhytidiadelphus
|family=Hylocomiaceae
|distribution=North America;Eurasia;Atlantic Islands;Pacific Islands (New Zealand);Australia (Tasmania);cool temperate and boreal zones
|reference=None
|publication title=Krypt.-Fl. Brandenburg
|publication year=1906
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_514.xml
|genus=Rhytidiadelphus
|apex shape=acuminate
|base shape=decurrent
|branch leaf shape=ovate;narrowly lanceolate
|capsule orientation=horizontal
|cell height or length or size=shorter
|cell variability=not differentiated
|cell width=wider
|costa quantity=0;2/3
|endostome segment architecture=perforate
|exostome tooth architecture or coloration or relief=reticulate
|exostome tooth pubescence=cross-striolate
|laminal cell architecture or pubescence or relief=coarsely;smooth
|leaf-stem architecture or arrangement or shape or vernation=plicate
|leaf-stem arrangement=not;crowded;falcate-secund
|leaf-stem distance=2.1mm;4.9mm
|leaf-stem orientation=squarrose-recurved;erect-spreading;squarrose
|leaf-stem relief=rugose
|leaf-stem shape=ovatelanceolate;ovate-triangular;ovatelanceolate;ovate-triangular;ovate
|margin architecture or shape=serrulate;serrulate;nearly entire
|operculum shape=conic
|paraphyllium presence=absent
|stem architecture=leafy;remotely branched;subpinnate
|stem orientation=creeping;ascending or erect
|stem width=2mm;8mm
}}<!--

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