View source for Thelia

{{Treatment/ID
|accepted_name=Thelia
|accepted_authority=Sullivant in A. Gray
|publications={{Treatment/Publication
|title=in A. Gray, Manual ed.
|place=2, 660. 1856
|year=1856
}}
|basionyms=
|synonyms=
|hierarchy=Theliaceae;Thelia
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Theliaceae]]</div><div class="higher-taxon"><small>genus</small>[[Thelia]]</div></div>
|etymology=Greek thele, nipple, alluding to prominent leaf papillae
|volume=Volume 28
|mention_page=page 637, 639, 648
|treatment_page=page 638
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism growth form"><b>Plants </b>small to medium-sized.</span> <span class="statement" id="st-d0_s1" data-properties="stem architecture;stem architecture;branch orientation;branch architecture;branch architecture"><b>Stems </b>1-pinnate or irregularly branched, branches erect-ascending, simple or irregularly branched;</span> <span class="statement" id="st-d0_s2" data-properties="epidermal-cell shape">epidermal-cells elongate;</span> <span class="statement" id="st-d0_s3" data-properties="rhizoid coloration;rhizoid architecture;rhizoid architecture or pubescence or relief">rhizoids reddish-brown, densely branched, smooth.</span> <span class="statement" id="st-d0_s4" data-properties="leaf shape"><b>Leaves </b>deltoid-ovate;</span> <span class="statement" id="st-d0_s5" data-properties="base shape">base decurrent;</span> <span class="statement" id="st-d0_s6" data-properties="apex shape;apex condition or size;acumen length or size;acumen shape;acumen shape">apex broadly acute to obtuse, abruptly contracted, acumen long, apiculate or piliferous;</span> <span class="statement" id="st-d0_s7" data-properties="costa quantity;costa quantity;costa quantity">costa single or sometimes double, 1/2–3/4 leaf length, sometimes spurred or 2-fid;</span> <span class="statement" id="st-d0_s8" data-properties="leaf length;leaf length;cell shape;cell shape">alar cells quadrate or subrectangular;</span> <span class="statement" id="st-d0_s9" data-properties="laminal cell shape;laminal cell relief;papilla shape;papilla architecture;papilla architecture">laminal cells rhombic to fusiform, stoutly 1-papillose abaxially, papillae columnar, simple or 3–6-branched.</span> <span class="statement" id="st-d0_s10" data-properties="seta coloration;seta coloration;seta course;seta architecture or pubescence or relief"><b>Seta </b>yellowish when young, red when mature, flexuose when dry, smooth.</span> <span class="statement" id="st-d0_s11" data-properties="capsule shape;capsule architecture or shape;capsule course;capsule relief"><b>Capsule </b>cylindric to ovoid-cylindric, symmetric to somewhat curved, smooth to weakly wrinkled when dry;</span> <span class="statement" id="st-d0_s12" data-properties="mouth shape;proximal cell shape;proximal cell size;proximal cell coloration;wall width">exothecial cells at mouth oblate to quadrate, proximal cells quadrate to short-rectangular, enlarged, red, walls thin to thick;</span> <span class="statement" id="st-d0_s13" data-properties="">stomata on neck;</span> <span class="statement" id="st-d0_s14" data-properties="annulus prominence;annulus duration;annulus architecture or arrangement;annulus architecture or arrangement;cell-wall width">annulus rudimentary, persistent, 2-seriate or 3-seriate, cell-walls thin;</span> <span class="statement" id="st-d0_s15" data-properties="operculum shape">operculum conic-subulate;</span> <span class="statement" id="st-d0_s16" data-properties="exostome tooth fusion;exostome tooth coloration;exostome tooth arrangement or course or shape;membrane height or length or size;membrane architecture or pubescence or relief;external surface architecture or pubescence or relief;external surface relief;trabecula size or width;median line size or width;internal surface relief;trabecula fragility">exostome teeth united at base on short, smooth membrane, white, linear, external surface smooth basally, finely papillose distally, trabeculae and median line thickened, internal surface lightly papillose, trabeculae weak;</span> <span class="statement" id="st-d0_s17" data-properties="endostome prominence;endostome coloration;endostome relief;basal membrane position;basal membrane height;segment size;cilium quantity;cilium quantity;cilium length;cilium presence">endostome rudimentary, white, lightly papillose, basal membrane low or high, segments reduced to stubs or to 1/2 exostome length, cilia 1 or 2, nublike or absent.</span> <span class="statement" id="st-d0_s18" data-properties="spore relief"><b>Spores </b>smooth to lightly papillose.</span><!--

-->{{Treatment/Body
|distribution=North America;ne Mexico;West Indies (Dominican Republic)
|discussion=<p>Species 3 (3 in the flora).</p><!--
--><p>Thelia is remarkable in its glaucous, bluish green color, thick, densely tomentose mats, and tumid to julaceous stems and branches. The stem leaves are distinctively deltoid-ovate with rhombic cells, simple or 3–6-branched columnar papillae, and frequently long-ciliate margins. The sporophyte has a long seta and typically erect, symmetric, cylindric to ovoid-cylindric capsules. The diplolepidous peristome is yellowish white. The endostome segments and cilia are often reduced to nublike projections on a low or high basal membrane.</p><!--
--><p>In the field, Thelia could be confused with Myurella (Pterigynandraceae) because both genera have terete-foliate stems and branches that occur in mats or cushions, radiculose stems, and a similar color. Myurella rarely occurs in dense mats, and its leaves have short, double costae, serrate or sinuate-dentate margins, and lower, unbranched papillae.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Stem leaf laminal cell papillae simple.
|[[Thelia hirtella|Thelia hirtella]]
|-id=key-0-1
|1
|Stem leaf laminal cell papillae usually 3-6-branched
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Stems creeping, densely and regularly 1-pinnate, often densely tomentose, branches simple; leaf acumina usually spreading-piliferous.
|[[Thelia asprella|Thelia asprella]]
|-id=key-0-2
|2
|Stems often ascending, sparsely and irregularly branched, lightly tomentose, branches irregularly branched; leaf acumina erect, apiculate.
|[[Thelia lescurii|Thelia lescurii]]
|}
</div></div><!--

-->{{#Taxon:
name=Thelia
|authority=Sullivant in A. Gray
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Theliaceae
|distribution=North America;ne Mexico;West Indies (Dominican Republic)
|reference=None
|publication title=in A. Gray, Manual ed.
|publication year=1856
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_1000.xml
|genus=Thelia
|acumen length or size=long
|acumen shape=piliferous;apiculate
|annulus architecture or arrangement=3-seriate;2-seriate
|annulus duration=persistent
|annulus prominence=rudimentary
|apex condition or size=contracted
|apex shape=broadly acute;obtuse
|basal membrane height=high
|basal membrane position=low
|base shape=decurrent
|branch architecture=branched;simple
|branch orientation=erect-ascending
|capsule architecture or shape=symmetric
|capsule course=curved
|capsule relief=smooth;weakly wrinkled
|capsule shape=cylindric;ovoid-cylindric
|cell shape=subrectangular;quadrate
|cell-wall width=thin
|cilium length=nublike
|cilium presence=absent
|cilium quantity=2;1
|costa quantity=1/2;3/4
|endostome coloration=white
|endostome prominence=rudimentary
|endostome relief=papillose
|epidermal-cell shape=elongate
|exostome tooth arrangement or course or shape=linear
|exostome tooth coloration=white
|exostome tooth fusion=united
|external surface architecture or pubescence or relief=smooth
|external surface relief=papillose
|internal surface relief=papillose
|laminal cell relief=1-papillose
|laminal cell shape=rhombic;fusiform
|leaf length=2-fid;spurred
|leaf shape=deltoid-ovate
|median line size or width=thickened
|membrane architecture or pubescence or relief=smooth
|membrane height or length or size=short
|mouth shape=oblate;quadrate
|operculum shape=conic-subulate
|papilla architecture=3-6-branched;simple
|papilla shape=columnar
|proximal cell coloration=red
|proximal cell shape=quadrate;short-rectangular
|proximal cell size=enlarged
|rhizoid architecture=branched
|rhizoid architecture or pubescence or relief=smooth
|rhizoid coloration=reddish-brown
|segment size=reduced
|seta architecture or pubescence or relief=smooth
|seta coloration=red;yellowish
|seta course=flexuose
|spore relief=smooth;lightly papillose
|stem architecture=branched;1-pinnate
|trabecula fragility=weak
|trabecula size or width=thickened
|wall width=thin;thick
|whole_organism growth form=plant
|whole_organism size=small;medium-sized
}}<!--

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