View source for Toiyabea ← Toiyabea You do not have permission to edit this page, for the following reason: The action you have requested is limited to users in the group: Users. You can view and copy the source of this page. {{Treatment/ID |accepted_name=Toiyabea |accepted_authority=R. P. Roberts |publications={{Treatment/Publication |title=Sida |place=21: 1652. 2005 |year=2005 }} |common_names=Alpine serpentweed |basionyms= |synonyms= |hierarchy=Asteraceae;Asteraceae tribe Astereae;Toiyabea |hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Asteraceae]]</div><div class="higher-taxon"><small>tribe</small>[[Asteraceae tribe Astereae]]</div><div class="higher-taxon"><small>genus</small>[[Toiyabea]]</div></div> |etymology=For Toiyabe Mountain Range, Nevada; traditional Shoshone, black mountains |volume=Volume 20 |mention_page=page 13, 173, 182 |treatment_page=page 172 }}<!-- --><span class="statement" id="st-d0_s0" data-properties="whole-organism some measurement;whole-organism duration;herbage pubescence"><b>Perennials,</b> 10–20 cm, herbage densely stipitate-glandular (with caudices or short rhizomes, bases woody).</span> <span class="statement" id="st-d0_s1" data-properties="stem atypical quantity;stem orientation;stem architecture"><b>Stems </b>(1–10) erect, usually simple.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Leaves </b>basal and cauline;</span> <span class="statement" id="st-d0_s3" data-properties="">alternate;</span> <span class="statement" id="st-d0_s4" data-properties="leaf position;leaf position;leaf arrangement;leaf architecture;proximal leaf architecture">usually sessile, proximal petiolate;</span> <span class="statement" id="st-d0_s5" data-properties="midnerve prominence;midnerve quantity;collateral nerve size;collateral nerve arrangement;collateral nerve prominence;collateral nerve shape;collateral nerve architecture or shape;collateral nerve coloration;collateral margin size;collateral margin arrangement;collateral margin prominence;collateral margin shape;collateral margin architecture or shape;collateral margin coloration;collateral face size;collateral face arrangement;collateral face prominence;collateral face shape;collateral face architecture or shape;collateral face coloration">blades with prominent midnerves plus 1–2 pairs of smaller, ± parallel, collateral nerves often evident, spatulate to obovate or oblanceolate, margins (distal 1/2) coarsely dentate to serrate, faces often gland-dotted.</span> <span class="statement" id="st-d0_s6" data-properties="head architecture or shape;head arrangement;head atypical quantity;array shape;array shape;array architecture"><b>Heads </b>discoid, usually borne singly, sometimes (2–5) in elongate or flat-topped cymiform arrays.</span> <span class="statement" id="st-d0_s7" data-properties="involucre shape;involucre length;involucre width"><b>Involucres </b>campanulate to hemispheric, (10–12 ×) 7–10 mm.</span> <span class="statement" id="st-d0_s8" data-properties="phyllarie quantity;phyllarie architecture;phyllarie size;phyllarie architecture;series quantity;margin pubescence;face pubescence"><b>Phyllaries </b>21–28 in 2–3 series, 3-nerved (flat), subequal, foliaceous, margins ciliolate, faces stipitate-glandular;</span> <span class="statement" id="st-d0_s9" data-properties="outer phyllarie orientation;outer phyllarie shape;inner phyllarie shape">outer slightly spreading, broadly ovate (apices obtuse, mucronulate), inner lanceolate-spatulate (apices acuminate-cuspidate).</span> <span class="statement" id="st-d0_s10" data-properties="receptacle shape;receptacle relief;receptacle architecture"><b>Receptacles </b>convex, pitted, epaleate.</span> <span class="statement" id="st-d0_s11" data-properties="ray-floret presence"><b>Ray-</b>florets 0.</span> <span class="statement" id="st-d0_s12" data-properties="disc-floret atypical quantity;disc-floret atypical quantity;disc-floret quantity;disc-floret reproduction;disc-floret reproduction"><b>Disc-</b>florets (29–) 35–50 (–55), bisexual, fertile;</span> <span class="statement" id="st-d0_s13" data-properties="corolla coloration;tube height or length or size;lobe quantity;lobe orientation;lobe shape">corollas golden yellow, tubes shorter than narrowly funnelform throats, lobes 5, spreading to recurved, lanceolate;</span> <span class="statement" id="st-d0_s14" data-properties="style-branch appendage shape">style-branch appendages narrowly lanceolate.</span> <span class="statement" id="st-d0_s15" data-properties="cypsela shape;cypsela shape;cypsela architecture;face pubescence"><b>Cypselae </b>(tan to reddish-brown) cylindric to fusiform, sometimes slightly compressed, 4–5-nerved, faces hairy;</span> <span class="statement" id="st-d0_s16" data-properties="bristle quantity;bristle coloration;bristle width;bristle architecture;bristle shape;series quantity">pappi persistent, of ca. 25, tan, fine, barbellate, apically attenuate bristles in 1 series.</span> <span class="statement" id="st-d0_s17" data-properties="pappus duration;x chromosome quantity">x = 9.</span><!-- -->{{Treatment/Body |distribution=w United States |discussion=<p>Species 1.</p><!-- --><p>Toiyabea was originally described as Haplopappus and later transferred to Tonestus. Investigations based on DNA sequence data (R. P. Roberts 2002; Roberts and L. E. Urbatsch 2004) failed to support the monophyly of Tonestus in the sense of G. L. Nesom and D. R. Morgan (1990). Evolutionary affinities of Toiyabea appear to be with Petradoria. Close affinity of Toiyabea alpina to Tonestus eximius and Lorandersonia peirsonii suggested by morphology is not supported thus far by sequence data.</p> |tables= |references= }}<!-- --><!-- -->{{#Taxon: name=Toiyabea |author=Lowell E. Urbatsch;Roland P. Roberts;Kurt M. Neubig |authority=R. P. Roberts |rank=genus |parent rank=tribe |synonyms= |basionyms= |family=Asteraceae |illustrator=Yevonn Wilson-Ramsey |illustration copyright=Flora of North America Association |distribution=w United States |reference=None |publication title=Sida |publication year=2005 |special status= |source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V19-20-21/V20_377.xml |tribe=Asteraceae tribe Astereae |genus=Toiyabea |array architecture=cymiform |array shape=flat-topped;elongate |bristle architecture=barbellate |bristle coloration=tan |bristle quantity=25 |bristle shape=attenuate |bristle width=fine |collateral face architecture or shape=coarsely dentate;serrate |collateral face arrangement=parallel |collateral face coloration=gland-dotted |collateral face prominence=evident |collateral face shape=spatulate;obovate or oblanceolate |collateral face size=smaller |collateral margin architecture or shape=coarsely dentate;serrate |collateral margin arrangement=parallel |collateral margin coloration=gland-dotted |collateral margin prominence=evident |collateral margin shape=spatulate;obovate or oblanceolate |collateral margin size=smaller |collateral nerve architecture or shape=coarsely dentate;serrate |collateral nerve arrangement=parallel |collateral nerve coloration=gland-dotted |collateral nerve prominence=evident |collateral nerve shape=spatulate;obovate or oblanceolate |collateral nerve size=smaller |corolla coloration=golden yellow |cypsela architecture=4-5-nerved |cypsela shape=compressed;cylindric;fusiform |disc-floret atypical quantity=50;55 |disc-floret quantity=35;50 |disc-floret reproduction=fertile;bisexual |face pubescence=hairy;stipitate-glandular |head architecture or shape=discoid |head arrangement=singly |head atypical quantity=2;5 |herbage pubescence=stipitate-glandular |inner phyllarie shape=lanceolate-spatulate |involucre length=[10mm;12mm |involucre shape=campanulate;hemispheric |involucre width=]7mm;10mm |leaf architecture=sessile |leaf arrangement=alternate |leaf position=cauline;basal |lobe orientation=spreading;recurved |lobe quantity=5 |lobe shape=lanceolate |margin pubescence=ciliolate |midnerve prominence=prominent |midnerve quantity=1;2 |outer phyllarie orientation=spreading |outer phyllarie shape=ovate |pappus duration=persistent |phyllarie architecture=foliaceous;3-nerved |phyllarie quantity=21;28 |phyllarie size=subequal |proximal leaf architecture=petiolate |ray-floret presence=absent |receptacle architecture=epaleate |receptacle relief=pitted |receptacle shape=convex |series quantity=1;2;3 |stem architecture=simple |stem atypical quantity=1;10 |stem orientation=erect |style-branch appendage shape=lanceolate |tube height or length or size=shorter |whole-organism duration=perennial |whole-organism some measurement=10cm;20cm |x chromosome quantity=9 }}<!-- -->[[Category:Treatment]][[Category:Asteraceae tribe Astereae]] Templates used on this page: Template:Asteraceae (view source) Template:Treatment/AuthorLink (view source) Template:Treatment/Body (view source) Template:Treatment/Body/Maps (view source) Template:Treatment/ID (view source) Template:Treatment/Publication (view source) Return to Toiyabea.