View source for Ulota

{{Treatment/ID
|accepted_name=Ulota
|accepted_authority=D. Mohr
|publications={{Treatment/Publication
|title=Ann. Bot. (König & Sims)
|place=2: 540. 1806
|year=1806
}}
|basionyms=
|synonyms=
|hierarchy=Orthotrichaceae;Ulota
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Orthotrichaceae]]</div><div class="higher-taxon"><small>genus</small>[[Ulota]]</div></div>
|etymology=Greek, oulotes, curliness, alluding to leaves of some species
|volume=Volume 28
|mention_page=page 38, 50, 52, 63, 73, 78, 664, 667
|treatment_page=page 72
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism size;whole_organism growth form;cushion architecture or fragility"><b>Plants </b>small to medium-sized, in tufts or loose cushions.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;branch orientation"><b>Stems </b>erect or rarely creeping, branches erect to ascending.</span> <span class="statement" id="st-d0_s2" data-properties="leaf course;leaf shape;leaf course;leaf shape;leaf orientation;leaf course;leaf shape;leaf shape;leaf shape;leaf shape;leaf shape;leaf relief"><b>Leaves </b>straight and not crisped to flexuose and crisped when dry, erect-spreading to spreading-flexuose when moist, lanceolate, oblong-lanceolate, or linear-lanceolate, not rugose;</span> <span class="statement" id="st-d0_s3" data-properties="margin architecture or shape">margins entire;</span> <span class="statement" id="st-d0_s4" data-properties="apex shape;apex shape;apex shape;apex shape;apex shape">apex acuminate, acute, or narrowly obtuse;</span> <span class="statement" id="st-d0_s5" data-properties="costa architecture">costa ending near apex or rarely excurrent;</span> <span class="statement" id="st-d0_s6" data-properties="basal laminal cell shape">basal laminal cells roundedquadrate to elongate-rectangular or linear;</span> <span class="statement" id="st-d0_s7" data-properties="distal cell shape;distal cell some measurement;distal cell architecture or pubescence or relief;distal cell relief;papilla quantity;papilla shape;papilla shape;papilla shape;papilla shape;papilla shape">distal cells hexagonal-rounded to elliptic, 6–14 µm, sometimes smooth, usually papillose only over lumina, papillae 1 (or 2) per cell, conic, 2-fid, or clavate;</span> <span class="statement" id="st-d0_s8" data-properties="">marginal cells abruptly shorter than basal.</span> <span class="statement" id="st-d0_s9" data-properties="marginal cell height or length or size;marginal cell reproduction;marginal cell reproduction;marginal cell reproduction"><b>Sexual </b>condition autoicous or rarely dioicous;</span> <span class="statement" id="st-d0_s10" data-properties="perichaetial leaf size">perichaetial leaves larger than stem-leaves.</span> <span class="statement" id="st-d0_s11" data-properties="seta some measurement"><b>Seta </b>1–10 mm.</span> <span class="statement" id="st-d0_s12" data-properties="capsule position;capsule shape;capsule shape;capsule shape;capsule shape;capsule shape;capsule architecture or arrangement or shape or vernation;mouth architecture or shape;mouth architecture or shape;mouth character;mouth architecture or pubescence or relief;mouth relief;capsule size"><b>Capsule </b>fully exserted, fusiform-cylindric, ovate-oblong, or rarely obovate, slightly 8-plicate at mouth to strongly 8-ribbed entire length or rarely smooth, mouth puckered or capsule ± constricted below mouth;</span> <span class="statement" id="st-d0_s13" data-properties="stoma position">stomata superficial;</span> <span class="statement" id="st-d0_s14" data-properties="peristome quantity">peristome double or single;</span> <span class="statement" id="st-d0_s15" data-properties="exostome tooth quantity;split quantity;split relief;split pubescence;split pubescence">exostome teeth 8, sometimes split to 16, finely and densely papillose to papillose-striate or rarely smooth;</span> <span class="statement" id="st-d0_s16" data-properties="endostome segment quantity;endostome segment presence">endostome segments 8 or absent.</span> <span class="statement" id="st-d0_s17" data-properties="calyptra shape;calyptra height or length or size;calyptra shape;split quantity;split architecture or pubescence or relief;split pubescence;split architecture or arrangement or shape or vernation;capsule quantity"><b>Calyptra </b>mitrate, short to oblong-conic, base usually deeply split several times, smooth, hairy, not plicate, covering 1/2 capsule.</span> <span class="statement" id="st-d0_s18" data-properties=""><b>Spores </b>isosporous.</span><!--

-->{{Treatment/Body
|distribution=North America;South America;Europe;Asia;Pacific Islands (New Zealand);Australia;temperate regions
|discussion=<p>Species ca. 60 (9 in the flora).</p><!--
--><p>Ecologically, Ulota is primarily found either in wet, shaded coniferous forests, particularly in areas of high rainfall, or in subalpine areas on small trees. Ulota is closely related to Orthotrichum. Ulota is distinguished from other genera by the conic calyptra, highly differentiated basal laminal cells, crisped leaves, superficial stomata, and lack of gemmae on the leaves. The leaf base is usually ovate and clasping the stem; the interior basal laminal cells radiate from the insertion and are thick-walled, often nodose, and usually orange; the capsule is gradually contracted to the seta with a long neck; the prostome is absent; the exostome teeth are more or less perforate at their apices; and the endostome segments are incurved. A few species show a close resemblance to Macromitrium in having creeping stems and similar basal cells and calyptrae. The North American species can be divided into four groups: dioicous species, producing gemmae (U. phyllantha); species with a single, erect-flexuose peristome and leaves not much crisped (U. coarctata and U. drummondii); species with a double peristome and exostome teeth reflexed (U. barclayi, U. crispa, U. curvifolia, U. hutchinsiae, and U. obtusiuscula); and species with creeping branched stems (U. megalospora).</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Sexual condition dioicous; specialized asexual reproduction present on leaf apices; maritime habitats.
|[[Ulota phyllantha|Ulota phyllantha]]
|-id=key-0-1
|1
|Sexual condition autoicous; specialized asexual reproduction absent; non-maritime habitats
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Capsules with mouth small, distinctly smaller at mouth than at middle; peristome single; exostome teeth erect, flexuose
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Capsules with mouth wide but constricted below mouth or evenly tapering to seta from mouth; peristome double; exostome teeth reflexed
|[[#key-0-4| > 4]]
|-id=key-0-3
|3
|Capsules obovate to oblong-ovoid, puckered and slightly 8-plicate at mouth.
|[[Ulota coarctata|Ulota coarctata]]
|-id=key-0-3
|3
|Capsules cylindric to fusiform, 8-ribbed 1/2 -2/3 length.
|[[Ulota drummondii|Ulota drummondii]]
|-id=key-0-4
|4
|Perichaetial leaves differentiated from stem leaves.
|[[Ulota barclayi|Ulota barclayi]]
|-id=key-0-4
|4
|Perichaetial leaves not differentiated from stem leaves
|[[#key-0-5| > 5]]
|-id=key-0-5
|5
|Stems both creeping and erect; leaf apices filiform-acuminate; leaves linear- lanceolate; stomata near mid capsule; spores 35-60 µm.
|[[Ulota megalospora|Ulota megalospora]]
|-id=key-0-5
|5
|Stems all erect; leaf apices acute or obtuse; leaves lanceolate or linear-lanceolate; stomata in capsule neck; spores 9-32 µm
|[[#key-0-6| > 6]]
|-id=key-0-6
|6
|Leaves erect-appressed, not crisped or contorted when dry.
|[[Ulota hutchinsiae|Ulota hutchinsiae]]
|-id=key-0-6
|6
|Leaves flexuose, crisped, or contorted when dry
|[[#key-0-7| > 7]]
|-id=key-0-7
|7
|Plants on rock; leaves flexuose, not contorted or twisted when dry.
|[[Ulota curvifolia|Ulota curvifolia]]
|-id=key-0-7
|7
|Plants on trees; leaves contorted, crisped, or twisted when dry
|[[#key-0-8| > 8]]
|-id=key-0-8
|8
|Basal laminal cell walls thin, cells hexagonal-rounded, not pigmented; calyptrae sparsely hairy; leaves slightly crisped and curved when dry.
|[[Ulota barclayi|Ulota barclayi]]
|-id=key-0-8
|8
|Basal laminal cell walls thick, cells linear-elongate grading to quadrate at margins, pale yellow; calyptrae very hairy; leaves crisped-contorted when dry
|[[#key-0-9| > 9]]
|-id=key-0-9
|9
|Plants to 2.5 cm; setae to 3 mm; capsules constricted below mouth; leaves crisped-contorted, not twisted when dry; e North America.
|[[Ulota crispa|Ulota crispa]]
|-id=key-0-9
|9
|Plants to 4 cm; setae usually 4 mm or longer; capsules evenly tapering from mouth to seta; leaves contorted, some tightly twisted when dry; w North America.
|[[Ulota obtusiuscula|Ulota obtusiuscula]]
|}
</div></div><!--

-->{{#Taxon:
name=Ulota
|authority=D. Mohr
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Orthotrichaceae
|distribution=North America;South America;Europe;Asia;Pacific Islands (New Zealand);Australia;temperate regions
|reference=None
|publication title=Ann. Bot. (König & Sims)
|publication year=1806
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V28/V28_110.xml
|genus=Ulota
|apex shape=obtuse;acute;obtuse;acute;acuminate
|basal laminal cell shape=roundedquadrate;elongate-rectangular or linear
|branch orientation=erect;ascending
|calyptra height or length or size=short
|calyptra shape=oblong-conic;mitrate
|capsule architecture or arrangement or shape or vernation=8-plicate
|capsule position=exserted
|capsule quantity=1/2
|capsule shape=obovate;ovate-oblong;obovate;ovate-oblong;fusiform-cylindric
|capsule size=constricted
|costa architecture=excurrent
|cushion architecture or fragility=loose
|distal cell architecture or pubescence or relief=smooth
|distal cell relief=papillose
|distal cell shape=hexagonal-rounded;elliptic
|distal cell some measurement=6um;14um
|endostome segment presence=absent
|endostome segment quantity=8
|exostome tooth quantity=8
|leaf course=spreading-flexuose;flexuose;straight
|leaf orientation=erect-spreading
|leaf relief=not rugose
|leaf shape=linear-lanceolate;oblong-lanceolate;linear-lanceolate;oblong-lanceolate;lanceolate;crisped;not crisped
|margin architecture or shape=entire
|marginal cell height or length or size=abruptly shorter
|marginal cell reproduction=dioicous;autoicous;sexual
|mouth architecture or pubescence or relief=smooth
|mouth architecture or shape=entire;8-ribbed
|mouth character=length
|mouth relief=puckered
|papilla quantity=1
|papilla shape=clavate;2-fid;clavate;2-fid;conic
|perichaetial leaf size=larger
|peristome quantity=single
|seta some measurement=1mm;10mm
|split architecture or arrangement or shape or vernation=not plicate
|split architecture or pubescence or relief=smooth
|split pubescence=hairy;smooth;papillose-striate
|split quantity=several;16
|split relief=papillose
|stem orientation=creeping;erect
|stoma position=superficial
|whole_organism growth form=plant
|whole_organism size=small;medium-sized
}}<!--

-->[[Category:Treatment]][[Category:Orthotrichaceae]]