View source for Wyethia

{{Treatment/ID
|accepted_name=Wyethia
|accepted_authority=Nuttall
|publications={{Treatment/Publication
|title=J. Acad. Nat. Sci. Philadelphia
|place=7: 39, plate 5. 1834
|year=1834
}}
|common_names=Mules-ears
|basionyms=
|synonyms=
|hierarchy=Asteraceae;Asteraceae tribe Heliantheae;Asteraceae (tribe Heliantheae) subtribe Ecliptinae;Wyethia
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Asteraceae]]</div><div class="higher-taxon"><small>tribe</small>[[Asteraceae tribe Heliantheae]]</div><div class="higher-taxon"><small>subtribe</small>[[Asteraceae (tribe Heliantheae) subtribe Ecliptinae]]</div><div class="higher-taxon"><small>genus</small>[[Wyethia]]</div></div>
|etymology=For Nathaniel Jarvis Wyeth, 1802–1856, early western American explorer
|volume=Volume 21
|mention_page=page 65, 66
|treatment_page=page 100
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole-organism atypical some measurement;whole-organism some measurement;whole-organism duration"><b>Perennials,</b> 15–60 (–100) cm (taproots relatively massive; caudices seldom branched).</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem architecture"><b>Stems </b>erect (or bending erect from bases), branched mostly from bases.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Leaves </b>basal and cauline;</span> <span class="statement" id="st-d0_s3" data-properties="">alternate;</span> <span class="statement" id="st-d0_s4" data-properties="leaf position;leaf position;leaf arrangement;leaf architecture;leaf architecture">usually petiolate (basal), sometimes sessile;</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;blade shape;blade arrangement or shape;blade shape;blade shape;blade shape;blade shape;base shape;margin architecture or shape;margin architecture or shape;margin shape or architecture;face pubescence;face pubescence">blades (usually pinnately nerved, basal larger than cauline) ± deltate, elliptic-ovate, lanceolate, lance-elliptic, lance-linear, or oblong-ovate, bases truncate to cuneate, margins usually entire, rarely dentate to serrate (sometimes ciliate), faces glabrous or hairy (sometimes glanddotted or finely stipitate-glandular).</span> <span class="statement" id="st-d0_s6" data-properties="head architecture or arrangement;head arrangement;head atypical quantity;head quantity;array architecture"><b>Heads </b>radiate, borne singly or 2–5 (–8+) in ± corymbiform to racemiform arrays.</span> <span class="statement" id="st-d0_s7" data-properties="involucre shape;involucre diameter"><b>Involucres </b>hemispheric to campanulate or turbinate, 12–60+ mm diam.</span> <span class="statement" id="st-d0_s8" data-properties="phyllary duration;phyllary atypical quantity;phyllary quantity;series quantity"><b>Phyllaries </b>persistent, 12–36 (–48) in 2–3+ series (subequal to unequal, outer sometimes foliaceous, much larger than inner).</span> <span class="statement" id="st-d0_s9" data-properties="receptacle shape;receptacle architecture"><b>Receptacles </b>flat to convex, paleate (paleae conduplicate, at least bases, papery).</span> <span class="statement" id="st-d0_s10" data-properties="ray-floret quantity;ray-floret architecture;ray-floret reproduction"><b>Ray-</b>florets 5–25+, pistillate, fertile;</span> <span class="statement" id="st-d0_s11" data-properties="corolla coloration">corollas mostly yellow (cream to white in W. helianthoides).</span> <span class="statement" id="st-d0_s12" data-properties="disc-floret quantity;disc-floret reproduction;disc-floret reproduction"><b>Disc-</b>florets 35–150+, bisexual, fertile;</span> <span class="statement" id="st-d0_s13" data-properties="corolla coloration;tube height or length or size;throat shape">corollas yellow, tubes much shorter than cylindric throats;</span> <span class="statement" id="st-d0_s14" data-properties="lobe quantity;lobe shape">lobes 5, ± deltate to lanceolate (style-branches stigmatic in 2 barely distinct lines, appendages ± filiform).</span> <span class="statement" id="st-d0_s15" data-properties="cypsela shape;cypsela shape"><b>Cypselae </b>± prismatic, weakly 3–4-angled (faces glabrous or hairy);</span> <span class="statement" id="st-d0_s16" data-properties="scale quantity;scale shape;scale architecture or relief;scale shape">pappi 0, or persistent, coroniform (usually lacerate) or of 1–4+ ovate to subulate, erose to lacerate (often basally connate) scales.</span> <span class="statement" id="st-d0_s17" data-properties="pappus presence;pappus duration;pappus shape;pappus shape;x chromosome quantity">x = 19.</span><!--

-->{{Treatment/Body
|distribution=w North America
|discussion=<p>Species 8 (8 in the flora).</p><!--
--><p>Some species formerly included in Wyethia are here treated in Agnorhiza and Scabrethia. Balsamorhiza is closely related to Wyethia.</p>
|tables=
|references={{Treatment/Reference
|id=weber1946a
|text=Weber, W. A. 1946. A taxonomic and cytological study of the genus Wyethia, family Compositae, with notes on the related genus Balsamorhiza. Amer. Midl. Naturalist 35: 400–452.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Involucres 35–60+ mm diam.; outer phyllaries (30–)40–80+ mm (± foliaceous, much surpassing discs)
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Involucres 10–30(–40) mm diam.; outer phyllaries 15–30(–40) mm (not foliaceous, seldom surpassing discs)
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Leaf blades: faces glabrous or finely stipitate-glandular, sometimes sparsely pilosulous as well (usually shining); cypselae 10–13 mm
|[[Wyethia glabra|Wyethia glabra]]
|-id=key-0-2
|2
|Leaf blades: faces densely tomentose to tomentulose (usually gland-dotted as well), glabrescent; cypselae 12–15 mm
|[[Wyethia helenioides|Wyethia helenioides]]
|-id=key-0-3
|3
|Leaf blades: faces tomentose to tomentulose (at least when young, glabrescent, usuallygland-dotted)
|[[Wyethia mollis|Wyethia mollis]]
|-id=key-0-3
|3
|Leaf blades: faces glabrous, glabrate, hirsute, hirsutulous, hirtellous, pilose, pilosulous, scabrellous, or strigillose (not tomentose to tomentulose, often gland-dotted or stipitate-glandular)
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Leaf blades: faces glabrous or glabrate (gland-dotted); phyllary margins not ciliate; cypselae glabrous
|[[#key-0-5| > 5]]
|-id=key-0-4
|4
|Leaf blades: faces usually hirsute, hirsutulous, hirtellous, pilose, pilosulous, scabrellous, or strigillose (sometimes vernicose, rarely finely stipitate-glandular or gland-dotted), sometimes glabrate; phyllary margins ± ciliate; cypselae strigillose or glabrous
|[[#key-0-6| > 6]]
|-id=key-0-5
|5
|Basal leaves: blades lance-elliptic or oblong-lanceolate to lanceolate; rays 8–21(–25), laminae 25–60 mm; cypselae 8–9 mm
|[[Wyethia amplexicaulis|Wyethia amplexicaulis]]
|-id=key-0-5
|5
|Basal leaves: blades narrowly oblong-lanceolate to lance-linear; rays 5–8+, lami-nae 18–30 mm; cypselae 6–8 mm
|[[Wyethia longicaulis|Wyethia longicaulis]]
|-id=key-0-6
|6
|Basal leaves: blades lanceolate to lance-linear; cypselae 7–8 mm, ± strigillose
|[[Wyethia angustifolia|Wyethia angustifolia]]
|-id=key-0-6
|6
|Basal leaves: blades elliptic-ovate, lanceolate, or lance-elliptic; cypselae 9–11 mm, strigillose or glabrous
|[[#key-0-7| > 7]]
|-id=key-0-7
|7
|Phyllaries 36–48; rays 13–25, laminae (cream to white) 25–45 mm; cypselae strigillose distally and on margins
|[[Wyethia helianthoides|Wyethia helianthoides]]
|-id=key-0-7
|7
|Phyllaries 16–34; rays 11–12, laminae (yellow) 25–50 mm; cypselaeglabrous
|[[Wyethia arizonica|Wyethia arizonica]]
|}
</div></div><!--

-->{{#Taxon:
name=Wyethia
|author=William A. Weber
|authority=Nuttall
|rank=genus
|parent rank=subtribe
|synonyms=
|basionyms=
|family=Asteraceae
|illustrator=John Myers
|illustration copyright=Flora of North America Association
|distribution=w North America
|reference=weber1946a
|publication title=J. Acad. Nat. Sci. Philadelphia
|publication year=1834
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V19-20-21/V21_237.xml
|tribe=Asteraceae tribe Heliantheae
|subtribe=Asteraceae (tribe Heliantheae) subtribe Ecliptinae
|genus=Wyethia
|array architecture=less corymbiform;racemiform
|base shape=truncate;cuneate
|blade arrangement or shape=lance-elliptic
|blade shape=oblong-ovate;lance-linear;oblong-ovate;lance-linear;lanceolate;elliptic-ovate;deltate
|corolla coloration=yellow;yellow
|cypsela shape=3-4-angled;prismatic
|disc-floret quantity=35;150
|disc-floret reproduction=fertile;bisexual
|face pubescence=hairy;glabrous
|head architecture or arrangement=radiate
|head arrangement=singly
|head atypical quantity=5;8
|head quantity=2;5
|involucre diameter=12mm;60mm
|involucre shape=hemispheric;campanulate or turbinate
|leaf architecture=sessile;petiolate
|leaf arrangement=alternate
|leaf position=cauline;basal
|lobe quantity=5
|lobe shape=less deltate;lanceolate
|margin architecture or shape=dentate;entire
|margin shape or architecture=dentate to serrate
|pappus duration=persistent
|pappus presence=absent
|pappus shape=of 1-4+ ovate to subulate , erose to lacerate scales;coroniform
|phyllary atypical quantity=36;48
|phyllary duration=persistent
|phyllary quantity=12;36
|ray-floret architecture=pistillate
|ray-floret quantity=5;25
|ray-floret reproduction=fertile
|receptacle architecture=paleate
|receptacle shape=flat;convex
|scale architecture or relief=erose
|scale quantity=1;4
|scale shape=lacerate;ovate;subulate
|series quantity=2;3
|stem architecture=branched
|stem orientation=erect
|throat shape=cylindric
|tube height or length or size=much shorter
|whole-organism atypical some measurement=60cm;100cm
|whole-organism duration=perennial
|whole-organism some measurement=15cm;60cm
|x chromosome quantity=19
}}<!--

-->[[Category:Treatment]][[Category:Asteraceae (tribe Heliantheae) subtribe Ecliptinae]]