Brachystigma
Proc. Acad. Nat. Sci. Philadelphia 80: 432. 1928.
Herbs, perennial; hemiparasitic, caudex woody. Stems erect, not fleshy, hirsutulous. Leaves cauline, in whorls of 3; petiole absent; blade not fleshy, not leathery, margins entire. Inflorescences terminal, racemes; bracts present. Pedicels present; bracteoles absent. Flowers: sepals 5, calyx nearly radially symmetric, broadly campanulate, lobes deltate; petals 5, corolla yellow, bilabiate, subrotate, abaxial lobes 3, adaxial 2; stamens 4, didynamous, filaments glabrescent proximally, villous distally; staminode 0; ovary 2-locular, placentation axile; stigma clavate. Capsules: dehiscence loculicidal. Seeds 100+, dark-brown, ellipsoid, wings absent or present.
Distribution
sw United States, n Mexico
Discussion
Species 1.
Brachystigma is monospecific and narrowly restricted to dry mountain slopes of southeastern Arizona, southwestern New Mexico, and northern Mexico. F. W. Pennell (1928) differentiated it from the similar genus Agalinis by its yellow corollas, glabrous anthers, capitate stigmas, more acute capsules, and winged seeds. Phylogenetic analysis of three chloroplast genes supports this distinction (M. C. Neel and M. P. Cummings 2004). Further, Brachystigma may be differentiated from closely related genera Aureolaria, Dasistoma, and Seymeria by its leaves arranged in whorls of three, a characteristic unique in Orobanchaceae.