Oenothera psammophila

(A. Nelson & J. F. Macbride) W. L. Wagner, Stockhouse & W. M. Klein

Monogr. Syst. Bot. Missouri Bot. Gard. 12: 84. 1985.

Common names: St. Anthony Dunes evening primrose
Basionym: Pachylophus psammophilusa. Nelson & J. F. Macbride Bot. Gaz. 61: 32. 1916
Synonyms: Oenothera cespitosa var. psammophila (A. Nelson & J. F. Macbride) Munz
Treatment appears in FNA Volume 10.
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Herbs perennial, caulescent, glabrous, also with resinous exudate, especially on younger leaves; from woody taproot. Stems decumbent, usually branched, 10–30 cm, becoming woody and buried in sand. Leaves (6–) 8–9 (–14.2) × (0.7–) 1.5–2 (–3.1) cm; petiole 3–9 cm; blade narrowly oblanceolate to oblanceolate, margins usually subentire or repand or remotely dentate, rarely serrate, apex acute. Flowers 1–3 per stem opening per day near sunset, with sweet scent; buds erect; floral-tube 42–60 mm; sepals 22–28 mm; petals white, fading rose-pink to rose, 23–40 mm; filaments 17–19 mm, anthers 13–16 mm; style 60–72 (–88) mm, stigma exserted beyond anthers at anthesis. Capsules somewhat curved and often somewhat twisted, lanceoloid-cylindrical, nearly cylindrical, (20–) 30–47 × 7–8 mm, gradually tapering to apex, 6–8 mm, dehiscent nearly throughout their length, valve margins with a conspicuous, irregular, wavy ridge; pedicel 1–5 mm. Seeds numerous, in 2 distinct rows per locule, narrowly obovoid, 2.5–3 × 1.2–1.4 mm, embryo 7/8 of seed volume, surface finely reticulate; seed collar with a broad membrane sealing cavity, margin entire. 2n = 14.


Phenology: Flowering May–Jul.
Habitat: On barren areas of drifting sand at interface between outcrops of lava and sand dunes.
Elevation: 1500–1700 m.

Discussion

Oenothera psammophila is known only from the dune area north and west of St. Anthony, Fremont County. It is unique in Oenothera because of the exu­date produced on the leaves to which sand particles adhere, forming a sand sheath, presumably for protection from the constantly blowing sand particles. It is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005).

Selected References

None.

Lower Taxa

None.

No values specified.

... more about "Oenothera psammophila"
1.3 cm13 mm <br />0.013 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
Warren L. Wagner +
(A. Nelson & J. F. Macbride) W. L. Wagner, Stockhouse & W. M. Klein +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Pachylophus psammophilusa. +
indehiscent +  and loculicidal +
subentire +, serrate +, usually coarsely dentate +  and pinnatifid +
narrowly oblanceolate +  and oblanceolate +
quadrangular +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
indehiscent +  and dehiscent +
tapering +, cylindrical +  and lanceoloid-cylindrical +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
St. Anthony Dunes evening primrose +
1500–1700 m. +
whitish +  and green +
exfoliating +
basifixed +  and versatile +
1.7 cm17 mm <br />0.017 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
14 cm140 mm <br />0.14 m <br /> (16.5 cm165 mm <br />0.165 m <br />) +
hispid +, lanate +  and glabrous +
4.2 cm42 mm <br />0.042 m <br /> (6 cm60 mm <br />0.06 m <br />) +
zygomorphic +  and actinomorphic +
unscented +
1 +  and 3 +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
On barren areas of drifting sand at interface between outcrops of lava and sand dunes. +
perennial +
suffrutescent +
nodding +  and erect +
9 cm90 mm <br />0.09 m <br /> (14.2 cm142 mm <br />0.142 m <br />) +
26 cm260 mm <br />0.26 m <br /> (36 cm360 mm <br />0.36 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
deciduous +
8 cm80 mm <br />0.08 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (26 cm260 mm <br />0.26 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
entire +, serrate +  and dentate +
repand +  and subentire +
pedicellate +  and sessile +
1 +  and 8 +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
fading rose-pink +  and rose +
obcordate +  and obovate +
2.3 cm23 mm <br />0.023 m <br /> (4 cm40 mm <br />0.04 m <br />) +
3 cm30 mm <br />0.03 m <br /> (9 cm90 mm <br />0.09 m <br />) +
Flowering May–Jul. +
Monogr. Syst. Bot. Missouri Bot. Gard. +
conspicuous +
basal +  and cauline +
distinct +
clavate +  and globose +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
sculptured +  and smooth +
obovoid +
reduced +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
in pairs +  and separating +
purple +, pink +, red +  and green +
deciduous +
2.2 cm22 mm <br />0.022 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
decumbent +, erect +  and ascending +
10 cm100 mm <br />0.1 m <br /> (30 cm300 mm <br />0.3 m <br />) +
petiolate +  and sessile +
intrapetiolar +
7.2 cm72 mm <br />0.072 m <br /> (8.8 cm88 mm <br />0.088 m <br />) +
pubescent +  and glabrous +
6 cm60 mm <br />0.06 m <br /> (7.2 cm72 mm <br />0.072 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
roughened +, reticulate +  and papillose +
Oenothera cespitosa var. psammophila +
Oenothera psammophila +
Oenothera sect. Pachylophus +
species +
3(-5)-aperturate +
pedicellate +  and sessile +
woody +  and hard +
smooth +  and erose +
papillate +