Epilobium obscurum
Spic. Fl. Lips., 147, [155]. 1771.
Herbs with elongated, leafy epigeal stolons. Stems erect or ascending, subterete, 20–80 cm, often well branched from base, sometimes also distally, subglabrous proximal to inflorescence with raised strigillose lines decurrent from margins of petioles, strigillose distally. Leaves opposite proximal to inflorescence, alternate distally, petiole 0–2 mm; blade green or slightly bluish green, lanceolate to narrowly ovate, 1.5–10 × 0.4–1.8 cm, ± shorter than internodes, base rounded to attenuate, margins denticulate with 15–40 evenly spaced teeth per side, veins prominent, 3–7 per side, apex subacute, surfaces sparsely strigillose, mainly on margins and veins; bracts gradually reduced. Inflorescences erect racemes or sparse panicles, strigillose. Flowers erect; buds 2–5 × 1–2.5 mm; pedicel 2–14 mm; floral-tube 0.8–1 × 1.1–1.5 mm, conspicuous ring of spreading hairs at mouth inside, mixed strigillose and sparse glandular puberulent externally; sepals lanceolate, somewhat keeled, 2.5–4 × 1–1.3 mm, abaxial surface strigillose; petals rose-purple, 3.5–6 × 1.8–3 mm, apical notch 0.8–1.4 mm; filaments pale-pink, those of longer stamens 2–2.2 mm, those of shorter ones 0.8–1.3 mm; anthers yellow, 0.7–0.8 × 0.4–0.5 mm; ovary 12–38 mm, strigillose; style white, 2.5–3.5 mm, glabrous, stigma clavate, 1.5–2 × 0.6–0.8 mm, surrounded by longer anthers. Capsules 40–70 mm, surfaces strigillose; pedicel 4–16 mm. Seeds narrowly obovoid, 0.9–1 × 0.3–0.4 mm, chalazal collar inconspicuous, brown, surface coarsely papillose; coma readily detached, dull white, 4–5 mm. 2n = 36.
Phenology: Flowering Jul–Aug.
Habitat: Ruderal areas, banks of ditches, streams, edges of swampy areas.
Elevation: 0–200[–500] m.
Distribution
Introduced; B.C., Mich., Wash., Europe, also in South America (Chile), Africa (Morocco), Pacific Islands (New Zealand), Australia (Tasmania)
Discussion
Epilobium obscurum, native throughout Europe and the European part of Russia except the far north, to Turkey and the Azores, is one of several Eurasian species that has naturalized in North America, following multiple early introductions around east coast port cities, and later around the Great Lakes. E. G. Voss(1972–1996, vol. 2) reported a collection of E. obscurum made in 1927 in Michigan near Detroit, but despite efforts by Voss and others, no additional collections of this species have been detected in that area, suggesting that it failed to become naturalized there.
In the Pacific Northwest, W. Suksdorf grew and collected at least six European taxa in his garden in Bingen (Klickitat County, Washington); this included E. obscurum (as early as 1922) but none of those taxa became naturalized. However, one or more independent new introductions in the Seattle (P. Zika, pers. comm.) and Vancouver (F. Lomer, pers. comm.) regions appear to be more persistent and the species should be considered naturalized there.
Selected References
None.