Polygala smallii

R. R. Smith & D. B. Ward

Sida 6: 307. 1976.

Common names: Tiny or Small’s milkwort
Basionym: Polygalaarenicola small Bull. New York Bot. Gard. 3: 426. 1905
Synonyms: Pilostaxis arenicola Small
Treatment appears in FNA Volume 10.
Revision as of 20:03, 7 June 2022 by imported>Volume Importer
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Herbs biennial, usually single-stemmed, rarely multistemmed, 0.2–0.5 (–0.8) dm, unbranched; from taproot or fibrous-root cluster. Stems erect, glabrous. Leaves with persistent basal rosette, clustered and irregular, crowded; alternate; sessile or subsessile with narrowed petiolelike region to 5 (–10) mm (usually obscured by tightly clustered leaves); blade oblanceolate to linear-oblanceolate, 10–42 × 2–14 mm, succulent, base cuneate, apex usually rounded to obtuse, rarely acute, sometimes apiculate, surfaces glabrous. Racemes capitate, 0.4–3 × 0.5–1.8 cm; peduncle 0.5–5 cm; bracts deciduous, often tardily so, linear-subulate. Pedicels winged, 0.4–0.8 mm, glabrous. Flowers green or greenish yellow, 4.5–6 (–8) mm; sepals decurrent on pedicel, narrowly ovate to lanceolate, (2–) 3–5 mm, sometimes ciliolate; wings usually lanceolate, sometimes oblong-lanceolate, 4–6 (–8) × 1–2 mm, apex long-acuminate, sometimes slightly involute, tip 0.5–0.9 mm; keel (3.5–) 4 (–5.5) mm, crest 2-parted, with 3 2-fid or entire lobes on each side; stamens 6–8. Capsules broadly ellipsoid to subglobose, 1.6–2 × 1.2–1.6 mm, margins not winged. Seeds 1.9–2.3 mm, pubescent; aril 0.9–1.6 mm, lobes 1/2 to ± equal length of seed. 2n = 64, 68.


Phenology: Flowering spring (year-round).
Habitat: Pinelands.
Elevation: 0–50 m.

Discussion

Polygala smallii is known from Broward, Martin, Miami-Dade, Palm Beach, and St. Lucie counties.

Compared to Polygala nana, the most similar species, P. smallii is smaller and more compact (stems and inflorescences are shorter and almost always surpassed by the tightly clustered leaves), leaves are narrower with a less conspicuously narrowed petiole­like portion (less obviously spatulate), wing sepals are more strongly lanceolate (tapering more to the apex, whereas P. nana wings typically are more elliptic or oblong-lanceolate), flowers are much greener at anthesis, and seeds are longer.

Polygala smallii is in the Center for Plant Conser­vation’s National Collection of Endangered Plants.

Selected References

None.

Lower Taxa

None.
... more about "Polygala smallii"
long-acuminate +, acute +, usually rounded +  and obtuse +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
J. Richard Abbott +
R. R. Smith & D. B. Ward +
cuneate +
Polygalaarenicola small +
1 cm10 mm <br />0.01 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
oblanceolate +  and linear-oblanceolate +
uniform +
pubescent +  and glabrous +
succulent +
0.2 cm2 mm <br />0.002 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
deciduous +
linear-subulate +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
broadly ellipsoid +  and subglobose +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
Tiny or Small’s milkwort +
reduced +
2-parted +
0–50 m. +
sessile +  and pedicellate +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
greenish yellow +  and green +
in proximal leaf-axils +  and subterranean +
cleistogamous +  and chasmogamous +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
samaroid +  and pseudomonomerous +
glabrous +  and pubescent +
isodiametric +  and slightly oblong +
Pinelands. +
unbranched +  and single-stemmed +
0.4 cm4 mm <br />0.004 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
crested +
4 cm40 mm <br />0.04 m <br /> (?) +
subsessile +  and sessile +
alternate +  and crowded +
cauline +  and basal +
entire +  and 3-2-fid +
leaf-opposed +, axillary +  and terminal +
glabrous +, ciliolate +  and ciliate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Flowering spring (year-round). +
syncarpous +  and 2[-8]-carpellate +
0.4 cm4 mm <br />0.004 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
petiolelike +  and narrowed +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
usually brownish +  and whitish +
exfoliating +
ellipsoid +
0.19 cm1.9 mm <br />0.0019 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
persistent +
narrowly ovate +  and lanceolate +
unequal +
0.3 cm3 mm <br />0.003 m <br /> (?) +  and 0.5 cm5 mm <br />0.005 m <br /> (?) +
6 +  and 8 +
prostrate +  and decumbent +
not +  and petiolate +
Pilostaxis arenicola +
Polygala smallii +
Polygala +
species +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
crested +  and lobed +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
persistent +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
oblong-lanceolate +  and lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
1mm;2mm +