Saussurea

de Candolle

Ann. Mus. Natl. Hist. Nat. 16: 156, 196, plates 10–13. 1810.

Common names: Saw-wort
Etymology: For Nicolas Théodore (1767–1845) and Horace Bénédict (1740–1799) de Saussure, Swiss naturalists
Treatment appears in FNA Volume 19. Treatment on page 165. Mentioned on page 58, 83.
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 TaxonIllustrator 
FNA19 P08 Cirsium scariosum var amer.jpegCirsium scariosum var. americanum
Cirsium vinaceum
Silybum marianum
Saussurea americana
John Myers
John Myers
John Myers
John Myers

Perennials, 5–120+ cm; herbage tomentose or glabrescent, not spiny. Stems erect or ascending, simple or branched. Leaves basal or cauline (sometimes cauline only), sessile or petiolate; blade margins entire or dentate to pinnately lobed, faces glabrous to densely tomentose, glandular or eglandular. Heads discoid, borne singly or in corymbiform arrays. Involucres ovoid to campanulate or ± turbinate. Phyllaries many in 3–5 (–10+) series, subequal to strongly unequal, appressed or not, ovate to lanceolate, margins entire, toothed, or lobed, apices obtuse or acute, appendaged or not, not spine-tipped. Receptacles flat or convex, epaleate, smooth, usually subulate-scaly, sometimes bristly or naked. Florets 10–20; corollas white to blue or purple, tubes slender, abruptly expanded to throats, lobes linear; anther bases short-tailed, apical appendages linear, acute; style-branches: fused portions with minutely hairy subterminal nodes, distinct portions oblong to linear, short-papillate. Cypselae oblong, ± angled, cylindric or 4–5-angled, ribs (when present) smooth or roughened, apices entire, glabrous or minutely glandular, attachment scars basal; pappi usually of 2 series, outer of readily falling, short bristles, inner persistent or falling as unit, of basally connate, usually longer, plumose bristles. x = 13, 14, 16, 17, 18, 19?.

Distribution

North America, Eurasia, 1 in Australia

Discussion

Species 300–400 (6 in the flora).

Saussurea is a notoriously difficult, largely Asiatic genus with species boundaries often indistinct.

Key

1 Outer and mid phyllaries with toothed or lobed appendages Saussurea amara
1 Outer and mid phyllaries entire, without appendages > 2
2 Proximal leaves ovate to lanceolate, usually more than 30 mm wide, bases broadly obtuse to truncate or cordate; plants 30–120 cm > 3
2 Proximal leaves linear to elliptic, 2–25 mm wide, bases acute to acuminate; plants 3–40 cm > 4
3 Cauline leaves usually more than 20, finely to ± coarsely serrate or dentate; s Alaska and Yukon to California, Idaho, and Montana Saussurea americana
3 Cauline leaves usually 15 or fewer, coarsely laciniate-dentate; nw Alaska Saussurea triangulata
4 Phyllaries subequal, linear to lanceolate; receptacles naked Saussurea nuda
4 Phyllaries strongly unequal, the outer ovate to lanceolate, conspicuously shorter than inner; receptacles scaly > 5
5 Tips of outer and mid phyllaries acute; Alaska and nw Canada Saussurea angustifolia
5 Tips of outer and mid phyllaries ± rounded; Rocky Mountains Saussurea weberi

"fine" is not a number.

... more about "Saussurea"
short-tailed +
not +  and appendaged +
entire +  and not spine-tipped +
glandular +  and glabrous +
acute;obtuse +
scarious +
indeterminate +  and determinate +
David J. Keil +
de Candolle +
decurrent +
Asteraceae tribe Cynareae +
compound +  and simple +
dentate to pinnately lobed +  and entire +
10-nerved or 20-nerved +  and rugose +
Saw-wort +
actinomorphic +
white +  and blue or purple +
4-5-angled +, cylindric +, angled +  and oblong +
fertile +  and bisexual +
North America +, Eurasia +  and 1 in Australia +
For Nicolas Théodore (1767–1845) and Horace Bénédict (1740–1799) de Saussure, Swiss naturalists +
eglandular +  and glandular +
glabrous +  and densely tomentose +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
in corymbiform arrays +  and singly +
indeterminate +
glabrescent +  and tomentose +
each +  and sessile +
ovoid;campanulate or more or less turbinate +
petiolate +  and sessile +
cauline +  and basal +
deltate +, usually narrowly triangular +  and more or less linear +
entire +  and dentate +
lobed +, toothed +  and entire +
2-carpellate +
persistent +
papillate +  and smooth +
distinct +  and fused +
short-papillate +
oblong;linear +
Ann. Mus. Natl. Hist. Nat. +
naked +  and epaleate +
convex;flat +
lipschitz1979a +
roughened +  and smooth +
exalbuminous +
subequal;strongly unequal +
branched +  and simple +
ascending +  and erect +
appendaged +  and truncate +
dilated +  and swollen +
enlarged +
Compositae +
Saussurea +
Asteraceae tribe Cardueae +
expanded +  and slender +
5 cm50 mm <br />0.05 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
tree +, vine +, shrub +  and subshrub +
19 +, 18 +, 17 +, 16 +, 14 +  and 13 +