Rayjacksonia
Amer. J. Bot. 83: 368. 1996.
| Taxon | Illustrator ⠉ | |
|---|---|---|
 | Xanthocephalum gymnospermoides Rayjacksonia phyllocephala Grindelia arizonica | Barbara Alongi Barbara Alongi Yevonn Wilson-Ramsey |
Annuals to perennials, 4–80 cm, herbage glabrous, prominently sessile to stipitate-glandular; taprooted. Stems erect to decumbent-ascending, branched basally (leafy throughout). Leaves mostly cauline; alternate; sessile to subpetiolate; blades 1-nerved (pinnately veined), linear to oblanceolate, margins coarsely serrate (teeth bristle-tipped), rarely entire (apices acute to obtuse). Heads radiate, in paniculiform or subcorymbo-cymiform arrays. (Peduncles sometimes cobwebby.) Involucres hemispheric, (7–16 ×) 9–30 mm. Phyllaries 20–40 in 4–5 series, appressed or tips spreading to reflexed, 1-nerved (flat or proximally convex), linear to lanceolate or ovatelanceolate, equal to subequal, white-indurate, margins of inner scarious (apices erect, loosely spreading, or recurved, green-herbaceous, acute to acuminate), faces glabrous or glandular. Receptacles shallowly convex, pitted, epaleate. Ray-florets 14–46, pistillate, fertile; corollas yellow. Disc-florets 25–130, bisexual, fertile; corollas yellow, tubes shorter than abruptly ampliate throats, lobes 5, erect, triangular; style-branch appendages deltate. Cypselae dimorphic, faces strigoso-sericeous; ray broadly ellipsoid to obovoid, 3-angled (thick-walled), 0–4-ribbed; disc broadly ellipsoid to clavate, ± radially compressed (thinner-walled), 5–9-ribbed; pappi persistent, of 30–40 brownish, unequal, barbellate, attenuate bristles in 3–4 series (about as long as disc corollas), outer thinner, terete, inner thick-flattened. x = 6.
Distribution
s, c, se United States, n Mexico
Discussion
Species 3 (3 in the flora).
R. L. Hartman (1990) informally recognized the three species of Rayjacksonia as an undescribed genus most closely related to Grindelia, Isocoma, Xanthocephalum, and other genera of Machaerantherinae with x = 6. The taxonomy was formalized by M. A. Lane and Hartman (1996). The Rayjacksonia species are coherent in vestiture, spinulose leaf teeth, heads borne singly, and rays with prominent, yellow laminae.
Selected References
Key
| 1 | Mid-cauline leaf blades linear to linear-oblanceolate, 1–3(–4) mm wide; involucres 4–7 × 10–15 mm; rays 14–19, corollas 6.5–9.5 mm | Rayjacksonia aurea |
| 1 | Mid-cauline leaf blades oblanceolate to oblong or oblong-lanceolate, (1–)4–15(–25) mm wide; involucres (4.5–)6–15 × 9–30 mm; rays (13–)17–38, corollas 6–19 mm | > 2 |
| 2 | Heads on short, leafless or bracteate peduncles, not surpassed by distal leaves; mid-cauline leaf blades (1–)4–15(–25) mm wide; phyllaries in 3–4 series, strongly unequal, apices broadly spreading to squarrose, ca. 0.9–1 mm wide | Rayjacksonia annua |
| 2 | Heads essentially sessile, often surpassed by distal leaves that continue to very base of heads; mid-cauline leaf blades (2.5–)4–13 mm wide; phyllaries in 3–4 series, subequal, apices erect to spreading, 1.3–1.7 mm wide | Rayjacksonia phyllocephala |
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