Heterotheca villosa var. sierrablancensis

Semple

Rev. Heterotheca Phyllotheca, 147, fig. 49. 1996.

Common names: White Mountains hairy goldenaster
Treatment appears in FNA Volume 20. Treatment on page 254. Mentioned on page 250.
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Stems ascending to erect, 22–34 (–46) cm, moderately to densely hispido-strigose, abundantly long-hirsute, sparsely to moderately stipitate-glandular. Distal cauline leaf-blades (perpendicular proximally, ascending distally) lanceolate to ovate-oblong, 13–23 × (3–) 4–6.5 mm, ± reduced distally, bases rounded, margins flat, apices broadly acute to obtuse, faces moderately to densely strigose (hairs 29–118/mm2), sparsely or usually moderately stipitate-glandular (glands 2–50/mm2). Heads (1–) 3–12 (–17) in congested, corymbiform to subumbelliform arrays. Peduncles 6–31 (–49) mm, moderately to densely hispido-strigose, moderately glandular; bracts 1–3, linear-lanceolate to linear-oblong, reduced distally, not foliar. Involucres narrowly campanulate (fresh), 5.5–7.8 (–8.7) mm. Phyllaries narrowly triangular-lanceolate, margins often reddish purple distally, faces moderately short-strigose, eglandular or sparsely stipitate-glandular. Ray-florets (10–) 13–20, laminae (8.5–) 9–12 (–14) mm (showy). 2n = 18.


Phenology: Flowering Sep–Oct.
Habitat: Open, disturbed areas, exposed, igneous rocky slopes, rocky granitic outcrops, road cuts, forests of Abies, Pinus ponderosa, and/or Pseudotsuga
Elevation: 2200–3500 m

Discussion

Of conservation concern.

Variety sierrablancensis is distinguished by its lanceolate to ovate-oblong, moderately strigose and sparsely to usually moderately stipitate-glandular leaves, its subumbelliform arrays, and its showy rays. It is close to var. nana, but usually has a denser indument and more often ovate than oblong leaves. Some plants from the White Mountains approach Heterotheca fulcrata var. fulcrata in leaf shape but lack the distinct, large leafy bracts of the latter. Less hairy plants with oblong leaves are similar vegetatively to var. nana, but usually have longer, showier rays.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
broadly acute;obtuse +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
corymbiform +  and subumbelliform +
indeterminate +  and determinate +
John C. Semple +
Semple +
rounded +
Amellus villosus +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
rugulose +  and muricate +
not foliar +
linear-lanceolate +  and linear-oblong +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
White Mountains hairy goldenaster +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
2200–3500 m +
stipitate-glandular +, short-strigose +, or +  and strigose +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Open, disturbed areas, exposed, igneous rocky slopes, rocky granitic outcrops, road cuts, forests of Abies, Pinus ponderosa, and/or Pseudotsuga +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.78 cm7.8 mm <br />0.0078 m <br /> (0.87 cm8.7 mm <br />0.0087 m <br />) +
campanulate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.78 cm7.8 mm <br />0.0078 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.9 cm9 mm <br />0.009 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
3.1 cm31 mm <br />0.031 m <br /> (4.9 cm49 mm <br />0.049 m <br />) +
hispido-strigose +
0.6 cm6 mm <br />0.006 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
Flowering Sep–Oct. +
triangular-lanceolate +
ovate-oblanceolate +
Rev. Heterotheca Phyllotheca, +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
Illustrated +  and Endemic +
34 cm340 mm <br />0.34 m <br /> (46 cm460 mm <br />0.46 m <br />) +
ascending +  and erect +
abundantly long-hirsute +  and sparsely moderately stipitate-glandular +
1 +  and 50 +
22 cm220 mm <br />0.22 m <br /> (34 cm340 mm <br />0.34 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis villosa +, Diplogon villosum +  and Diplopappus villosus +
Heterotheca villosa var. sierrablancensis +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +