Ptychostomum creberrimum

(Taylor) J. R. Spence & H. P. Ramsay

Phytologia 87: 23. 2005.

Illustrated
Basionym: Bryum creberrimum Taylor London J. Bot. 5: 54. 1846
Synonyms: B. lisae var. cuspidatum (Bruch & Schimper) Margadant
Treatment appears in FNA Volume 28. Treatment on page 159. Mentioned on page 131, 157, 160, 161, 163, 165.
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Plants in dense or open turfs, green or yellow-green. Stems 1–2 (–3) cm, comose, innovations comose or somewhat elongate and evenly foliate; often strongly radiculose. Leaves green, twisted to contorted when dry, ovatelanceolate, flat to weakly concave, (1–) 2–3 (–3.5) mm, somewhat enlarged toward stem apex; base weakly decurrent; margins revolute to mid leaf or beyond, limbidium strong, in 2 or 3 rows, yellowish; apex acuminate; costa long-excurrent, awn ± smooth; proximal laminal cells 3–4: 1, same width or sometimes wider than more distal cells; medial and distal cells rhomboidal, 14–22 µm wide, 3–4: 1, walls usually thin to firm, never oblique to costa. Specialized asexual reproduction absent. Sexual condition synoicous. Seta 1–2 (–3) cm. Capsule brown, elongate-pyriform, symmetric, 2–4 mm, mouth yellow; operculum conic, apiculate; peristome well developed; exostome teeth yellow basally, hyaline distally, lamellae usually straight mid-tooth, pores absent along mid line; endostome not adherent to exostome, basal membrane high, 1/2 exostome height, segments with broadly ovate perforations, cilia long, appendiculate. Spores 10–14 (–16) µm, finely papillose, pale brown-yellow or green.


Phenology: Capsules mature May–Sep.
Habitat: Damp to dry soil, soil over rock
Elevation: low to high elevations (0-3500 m)

Distribution

V28 251-distribution-map.gif

Greenland, Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., P.E.I., Que., Sask., Yukon, Alaska, Ariz., Ark., Calif., Colo., Conn., Del., D.C., Idaho, Ill., Ind., Iowa, Kans., Ky., Maine, Md., Mass., Mich., Minn., Mo., Mont., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.Dak., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wyo., s South America, Eurasia, Pacific Islands (New Zealand), Australia

Discussion

Ptychostomum creberrimum is one of the most widespread and common species in the genus, occurring throughout the Northern Hemisphere and disjunct in the Southern Hemisphere. The species can be distinguished from the closely related P. lonchocaulon by its strictly synoicous sexual condition and narrower yellowish leaf border. It is similar also to P. pallescens, which is autoicous and has larger spores and elongate-ovate segment perforations.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number."narrow" is not a number.

... more about "Ptychostomum creberrimum"
acuminate +
John R. Spence +
(Taylor) J. R. Spence & H. P. Ramsay +
denticulate +
reduced +
curved +  and straight +
decurrent +
Bryum creberrimum +
cucullate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
nutant +, inclined +  and suberect +
long-exserted +
elongate-pyriform +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
inflated +
not differentiated +
appendiculate +
long-excurrent +
subpercurrent +
rhomboidal;hexagonal +
Greenland +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Ark. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Idaho +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, Nev. +, N.H. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.Dak. +, Tenn. +, Tex. +, Utah +, Vt. +, Va. +, Wash. +, W.Va. +, Wyo. +, s South America +, Eurasia +, Pacific Islands (New Zealand) +  and Australia +
low to high elevations (0-3500 m) +
hyaline;pale-yellow +
not adherent +
hyaline +, brown +, orange +  and yellow +
hyaline +  and yellow +
Damp to dry soil, soil over rock +
not +  and differentiated +
narrowly ovatelanceolate;triangular +
polyoicous +, monoicous +, dioicous +  and sexual +
imbricate;variously contorted +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
erect;erect-spreading +
flat +  and weakly concave +
enlarged +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
2-stratose +  and 1-stratose +
revolute entire +  and denticulate +
1 (?) +, 3 (?) +  and 4 (?) +
short-rectangular +  and quadrate +
lateral +  and terminal +
lateral +  and terminal +
diplolepidous-alternate +
Capsules mature May–Sep. +
differentiated +
smooth;papillose +
not absent +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
straight +  and flexuose +
elongate +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Illustrated +
green +  and pale brown-yellow +
germinating +
variable +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
B. lisae var. cuspidatum +
Ptychostomum creberrimum +
Ptychostomum subg. Cladodium +
species +
lanceolate +
distinct +
not +  and porose +
straight +
specialized +
synoicous +, sexual +  and asexual +
sinuate +
incrassate +
yellow-green +  and green +
large;small to very +