Cardamine umbellata

Greene

Pittonia 3: 154. 1897.

Synonyms: Cardamine hirsuta subsp. kamtschatica (Regel) O. E. Schulz Cardamine kamtschatica (Regel) Piper Cardamine oligosperma var. kamtschatica (Regel) Detling Cardamine sylvatica var. kamtschatica Regel
Treatment appears in FNA Volume 7. Treatment on page 484. Mentioned on page 464, 467.
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Perennials; usually glabrous. Rhizomes often elongated, usually slender, rarely thickened, 1–2 (–5) mm diam., (not fleshy). Stems (simple or few to several from base), erect to ascending, (not flexuous), unbranched basally, sometimes branched distally, (0.3–) 0.8–2.5 (–3) dm. Basal leaves (sometimes withered by anthesis), rosulate, pinnately compound, (3 or) 5 or 7 (or 9) -foliolate, 2–5 (–9) cm, leaflets petiolulate or subsessile; lateral leaflets shortly petiolulate or subsessile, blade usually broadly ovate, rarely broadly obovate or orbicular, smaller than terminal, margins usually entire, rarely slightly 3 (or 5) -lobed or crenate; terminal leaflet subsessile, blade reniform or orbicular, 0.4–0.8 (–1.2) cm × 5–9 (–16) mm, margins entire or 3 (or 5) -lobed or crenate. Cauline leaves 3–5 (–7), 3–7 (or 9) -foliolate, petiolate, leaflets subsessile or sessile; base not auriculate; lateral leaflets: blade narrowly obovate, oblanceolate to linear, margins similar to terminal; terminal leaflet blade narrowly obovate, ovate, oblanceolate, lanceolate, oblong, margins usually entire, sometimes 3-lobed or crenate. Racemes ebracteate, (subumbellate, 2–8 (–14) -flowered, rachis usually 3–20 mm). Fruiting pedicels suberect to ascending, 3–8 (–10) mm. Flowers: sepals (greenish or purplish), oblong, 1–2 × 0.5–1 mm, lateral pair not saccate basally; petals white, narrowly obovate, 2.5–5 × 1–3 mm. Fruits linear, (torulose), (1.3–) 1.8–2.5 (–3) cm × 0.8–1.5 (–2) mm; (valves glabrous or sparsely pubescent); style 0.5–2 mm. Seeds brown, oblong, 1–1.5 × 0.8–1 mm. 2n = 32, 36, 48.


Phenology: Flowering Jun–Sep.
Habitat: Stream banks, tundra, alpine slopes, wetlands, damp, swampy and mossy areas, beach gravel and sand, alpine stream margins
Elevation: 0-1800 m

Distribution

V7 748-distribution-map.gif

Alta., B.C., N.W.T., Yukon, Alaska, Wash., e Asia (Russian Far East)

Discussion

Recent molecular data (J. Lihová et al. 2006) indicate that Cardamine umbellata, often treated as a variety of C. oligosperma, represents a distinct lineage more closely related to taxa from New Zealand; this does not exclude C. oligosperma as one of the possible parents of this polyploid.

Selected References

None.

Lower Taxa

None.

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... more about "Cardamine umbellata"
48 +, 36 +  and 32 +
linear +, oblong +  and ovate +
Ihsan A. Al-Shehbaz +, Karol Marhold +  and Judita Lihová +
Greene +
7-foliolate +  and compound +
5 cm50 mm <br />0.05 m <br /> (9 cm90 mm <br />0.09 m <br />) +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
not auriculate +
appendaged +  and unappendaged +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
compound +  and trifoliolate +
lobed +, 1-3-pinnatisect +, toothed +  and entire +
oblanceolate;linear +
smaller +
0.5 cm5 mm <br />0.005 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and 3-7-foliolate +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
Alta. +, B.C. +, N.W.T. +, Yukon +, Alaska +, Wash. +  and e Asia (Russian Far East) +
0-1800 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
lilac;purple;lilac;purple;pink;white +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
ascending +, spreading +  and erect +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
latiseptate +  and unsegmented +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1.8 cm18 mm <br />0.018 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
angustiseptate +, 4-angled +  and terete +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
latiseptate +  and sessile +
torulose +, lanceolate +, oblong +  and linear +
Stream banks, tundra, alpine slopes, wetlands, damp, swampy and mossy areas, beach gravel and sand, alpine stream margins +
semiannular +  and annular +
subsessile +  and petiolulate +
compound +, pinnately +, trifoliolate +  and simple +
alternate +, not +  and rosulate +
basal +  and cauline +
sessile +, subsessile +  and petiolulate +
connate +  and distinct +
decurrent +
crenate;3-lobed;crenate;3-lobed;entire;crenate;3-lobed;entire +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
2.5mm;5mm +
rudimentary +
obovate +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Jun–Sep. +
trinucleate +  and 3(-11)-colpate +
2-8(-14)-flowered;subumbellate;ebracteate +
reflexed +, ascending +  and erect +
stout +  and slender +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
suberect +  and ascending +
flattened +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not +  and mucilaginous +
winged +  and margined +
uniseriate +  and biseriate +
not +  and mucilaginous +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
persistent +  and caducous +
distinct +
erect +  and spreading or ascending +
reduced +
latiseptate +  and unsegmented +
angustiseptate +, 4-angled +  and terete +
latiseptate +  and unsegmented +
angustiseptate +, 4-angled +  and terete +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
prostrate +  and decumbent +
erect +  and ascending +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Cardamine hirsuta subsp. kamtschatica +, Cardamine kamtschatica +, Cardamine oligosperma var. kamtschatica +  and Cardamine sylvatica var. kamtschatica +
Cardamine umbellata +
Cardamine +
species +
scale-like +  and forked +
pubescent +, , +  and glabrous +
anastomosing +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +