Boechera pendulocarpa

(A. Nelson) Windham & Al-Shehbaz

Harvard Pap. Bot. 11: 77. 2006.

Basionym: Arabis pendulocarpa A. Nelson Bot. Gaz. 30: 192. 1900
Synonyms: Arabis holboellii var. pendulocarpa (A. Nelson) Rollins Boechera holboellii var. pendulocarpa (A. Nelson) N. W. Snow
Treatment appears in FNA Volume 7. Treatment on page 395. Mentioned on page 352, 396, 400.

Perennials; short to long-lived; sexual; caudex usually not woody. Stems usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–3 dm, densely pubescent proximally, trichomes simple and stalked, 2–4-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally. Basal leaves: blade narrowly oblanceolate, 1.5–4 (–5) mm wide, margins entire, often ciliate along petiole, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. Cauline leaves: 6–17, sometimes concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. Racemes 4–11-flowered, unbranched. Fruiting pedicels arched, gently recurved, 3–8 (–12) mm, glabrous or sparsely pubescent, trichomes appressed, branched. Flowers ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid. Fruits pendent, not appressed to rachis, not secund, usually straight, edges parallel, (2–) 2.5–3.8 cm × 1.5–2.2 mm; valves glabrous; ovules 66–92 per ovary; style 0.3–0.5 mm. Seeds sub-biseriate, 1–1.2 × 0.7–0.9 mm; wing distal, 0.05–0.1 mm wide. 2n = 14.


Phenology: Flowering Apr–Jul.
Habitat: Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows
Elevation: 1000-3300 m

Distribution

V7 565-distribution-map.gif

B.C., Yukon, Calif., Idaho, Mont., Nev., Oreg., Utah, Wyo.

Discussion

Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993), B. pendulocarpa is easily distinguished from that species by having simple and 2–4-rayed (versus 4–8-rayed) trichomes proximally on stems, cauline leaves without auricles, fruiting pedicels gently (versus sharply) recurved, and shorter (2–)2.5–3.8 (versus 3.5–6.5) cm, non-secund fruits. The two taxa have allopatric distributions, with B. pendulocarpa found in the mountains of western North America and B. holboellii apparently confined to Greenland. Recent use of the name A. (Boechera) exilis for this taxon (e.g., G. A. Mulligan 1996; R. D. Dorn 2001; N. H. Holmgren 2005b) is based on misinterpretation of the type (M. D. Windham and I. A. Al-Shehbaz 2006).

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera pendulocarpa"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(A. Nelson) Windham & Al-Shehbaz +
not +  and rosulate +
Arabis pendulocarpa +
appendaged +  and unappendaged +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate +
much smaller +  and reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
B.C. +, Yukon +, Calif. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +  and Wyo. +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
2.5 cm25 mm <br />0.025 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
1000-3300 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
straight +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
lavender +  and white +
rudimentary +
oblanceolate +  and spatulate +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Flowering Apr–Jul. +
trinucleate +  and 3(-11)-colpate +
ellipsoid +
Harvard Pap. Bot. +
unbranched;4-11-flowered +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
margined +, not winged +  and winged +
sub-biseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
orbicular +, oblong +  and flattened +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
pubescent +  and glabrous +
Arabis holboellii var. pendulocarpa +  and Boechera holboellii var. pendulocarpa +
Boechera pendulocarpa +
Boechera +
species +
branched +, short-stalked +, stalked +  and simple +
pubescent +  and glabrous +
0.008 cm0.08 mm <br />8.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
perennial +  and long-lived +
pubescent +  and glabrous +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.05mm;0.1mm +