Botrychium rugulosum

W. H. Wagner

Contr. Univ. Michigan Herb. 15: 315. 1982.

Common names: St. Lawrence grapefern botryche à limbe rugueux
Endemic
Synonyms: Botrychium multifidum (S.G. Gmelin) Ruprecht forma dentatum R.M. Tryon
Treatment appears in FNA Volume 2.

Trophophore stalk 2 to 15 cm, 1–2.5 times length of trophophore rachis; blade green, finely rugulose and convex distally, 2–4-pinnate, to 15 × 26 cm, somewhat herbaceous. Pinnae to 9 pairs, usually approximate, horizontal to ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, divided to tip. Pinnules obliquely and angularly trowel-shaped to spatulate, margins usually denticulate, apex acute, venation pinnate. Sporophores 2-pinnate, 1–2 times length of trophophore. 2n =90.


Phenology: Leaves green over winter, appearing in midspring.
Habitat: In open fields and secondary forests over wide range in vicinity of St. Lawrence Seaway
Elevation: 200–1000 m

Distribution

V2 363-distribution-map.gif

Ont., Que., Mich., Minn., N.Y., Vt., Wis.

Discussion

The name "rugulosum" refers to the tendency of the segments to become more or less wrinkled and convex. Botrychium rugulosum occurs with B. dissectum, B. multifidum, and rarely B. oneidense. It is often found in small stands of only 5–10 individuals, but some populations number over 100.

Selected References

None.

Lower Taxa

None.
... more about "Botrychium rugulosum"
Warren H. Wagner Jr. +  and Florence S. Wagner +
W. H. Wagner +
Undefined sect. Sceptridium +
horizontal +  and ascending +
simple +  and 5-pinnate +
0 cm0 mm <br />0 m <br /> (15 cm150 mm <br />0.15 m <br />) +
rugulose +
leathery +  and herbaceous +
0 cm0 mm <br />0 m <br /> (26 cm260 mm <br />0.26 m <br />) +
St. Lawrence grapefern +  and botryche à limbe rugueux +
Ont. +, Que. +, Mich. +, Minn. +, N.Y. +, Vt. +  and Wis. +
200–1000 m +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
In open fields and secondary forests over wide range in vicinity of St. Lawrence Seaway +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
of widely scattered , long , uniseriate hairs +
denticulate +
Leaves green over winter, appearing in midspring. +
spreading;ascending +
angularly trowel-shaped +  and spatulate +
epiphytic +  and terrestrial +
spore-bearing +  and laminate +
photosynthetic +
with long , uniseriate hairs +  and glabrous +
Contr. Univ. Michigan Herb. +
flattened +
not fleshy +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
with corky ridges +  and smooth +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
not proliferous +
sessile +  and short-stalked +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
thick-walled +  and trilete +
1-2 times length of trophophore +
represented by hairy rudiment +
flattened +
not fleshy +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
unbranched +  and simple +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
tuberculate +  and rugate +
Botrychium multifidum +
Botrychium rugulosum +
Botrychium sect. Sceptridium +
species +
arising +  and erect +
1-2.5 times length of trophophore rachis +
2 cm20 mm <br />0.02 m <br /> (15 cm150 mm <br />0.15 m <br />) +
anastomosing +
pinnate +  and fan +