Diphasiastrum complanatum

(Linnaeus) Holub

Preslia 47: 108. 1975.

Common names: Northern running-pine lycopode aplati
Illustrated
Basionym: Lycopodium complanatum Linnaeus Sp. Pl. 2: 1104. 1753
Synonyms: Lycopodium complanatum var. canadense Victorin
Treatment appears in FNA Volume 2.

Horizontal stems on substrate surface or shallowly buried in litter, 1.1–2.2 mm wide; leaves appressed, linear to narrowly lanceolate, 1.4–4 × 0.5–1.2 mm, apices acute. Upright shoots 8–44 cm, branching irregularly successively to 5 times; leaves on upright main-stem appressed with decurrent base, narrowly lanceolate, 1.2–3.2 × 0.5–1.1 mm, apex acute to acuminate. Branchlets flat in cross-section, narrowly bladelike, 1.8–4 mm wide, annual bud constrictions abrupt and conspicuous; upperside green, faintly shiny, flat; underside dull, pale, flat. Leaves on branchlets 4-ranked; upperside leaves appressed, linear-lanceolate, free portion of blades 0.7–2 × 0.5–1.2 mm; lateral leaves appressed, 2.6–7.3 × 0.8–2.1 mm; underside leaves weakly developed, appressed, narrowly deltate, 0.7–1.5 × 0.4–0.9 mm. Peduncles 1–2 on each upright shoot, 0.5–8.5 × 0.4–0.9 cm; leaves spirally arranged to nearly whorled, linear-lanceolate, 1.4–4.1 × 0.4–1 mm, apex acute to blunt. Stalks forked at uniform distances. Strobili 1–2 (–4), 8.3–32 × 2–3 mm, apex blunt, sterile tip absent. Sporophylls broadly deltate to nearly cordate, 2–3 × 2–2.4 mm, apex abruptly tapering. 2n = 46.


Habitat: Dry open coniferous or mixed forest alpine slopes
Elevation: 0–2000 m

Distribution

V2 574-distribution-map.gif

Greenland, St. Pierre and Miquelon, Alta., B.C., Man., N.B., Nfld. and Labr. (Nfld.), N.W.T., N.S., Ont., P.E.I., Que., Sask., Yukon, Alaska, Idaho, Maine, Mich., Minn., Mont., N.H., N.Y., Vt., Wash., Wis., Wyo., circumboreal

Discussion

Diphasiastrum complanatum forms a hybrid with D. digitatum that is seemingly uncommon and has never received a binomial designation. It is probably far more common than collections indicate, however. Superficially, the hybrid resembles both parents and is often confused with them. Collections are known from Ontario, Quebec, Connecticut, Maine, Michigan, Minnesota, New Hampshire, Vermont, and Wisconsin.

Selected References

None.

Lower Taxa

None.
... more about "Diphasiastrum complanatum"
tapering;blunt;acute;blunt +
Warren H. Wagner Jr. +  and Joseph M. Beitel +
(Linnaeus) Holub +
decurrent +
Lycopodium complanatum +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Northern running-pine +  and lycopode aplati +
Greenland +, St. Pierre and Miquelon +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, N.S. +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Idaho +, Maine +, Mich. +, Minn. +, Mont. +, N.H. +, N.Y. +, Vt. +, Wash. +, Wis. +, Wyo. +  and circumboreal +
0–2000 m +
photosynthetic +
carrot--shaped +
Dry open coniferous or mixed forest alpine slopes +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.73 cm7.3 mm <br />0.0073 m <br />) +
0.8mm +  and 2.1mm +
pseudomonopodial +, dichotomous +, branching +, branched +  and simple +
flat +  and round +
spirally arranged +  and nearly whorled +
remote;dense +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.41 cm4.1 mm <br />0.0041 m <br />) +
spreading;ascending;spreading;ascending +
linear-lanceolate +, lanceolate +, linear +  and narrowly lanceolate +
larger +
trimorphic +  and uniform +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
aerial +  and subterranean +
0.5 cm5 mm <br />0.005 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
emerging +
0-5 times +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
quadrate +  and flattened +
Illustrated +
plectostelic +  and actino +
axillary +  and adaxial +
thick-walled +  and trilete +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
2mm +  and 2.4mm +
protostelic +
1.1mm;2.2mm +
subterranean +
stalked +  and sessile +
modified +  and unmodified +
photosynthetic +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
small-grooved +
subterranean +
reticulate +, rugulate +  and pitted +
Lycopodium complanatum var. canadense +
Diphasiastrum complanatum +
Diphasiastrum +
species +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
linear-lanceolate +