Euphorbia geyeri

Engelmann

Boston J. Nat. Hist. 5: 260. 1845.

Common names: Geyer’s sandmat
Synonyms: Chamaesyce geyeri (Engelmann) Small
Treatment appears in FNA Volume 12. Treatment on page 268. Mentioned on page 258, 262.

Herbs, annual, with taproot. Stems prostrate or slightly ascending, loosely mat-forming, 4–25 (–45) cm, glabrous. Leaves opposite; stipules usually distinct, occasionally connate basally on lower side, usually deeply parted into 3 or more filiform segments, 0.7–1.5 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong to oblong-obovate or oblongelliptic, 4–12 × 2–6 mm, base slightly asymmetric, angled or rounded, with one side usually expanded into small, rounded auricle, margins entire, apex usually truncate, occasionally emarginate, abaxial surface pale grayish green, both surfaces glabrous; only midvein conspicuous or venation obscurely pinnate (larger leaves). Cyathia solitary or in small, cymose clusters at distal nodes; peduncle 1–2 mm. Involucre broadly campanulate, 1–1.5 × 0.7–0.9 mm, glabrous; glands 4, green to reddish, elliptic-oblong to nearly circular, slightly cupped to folded, 0.2–0.4 × 0.2–0.6 mm; appendages rudimentary to absent or white to reddish-tinged, usually rounded, sometimes pointed, (0–) 0.5–1 × (0–) 0.1–1.2 mm, distal margin entire or slightly toothed. Staminate flowers 5–20. Pistillate flowers: ovary glabrous; styles 0.2–0.6 mm, 2-fid nearly 1/2 length. Capsules globose-ovoid, 1.5–2 × 1.5–3 mm, glabrous; columella 1.5–1.9 mm. Seeds ashy white, ovoid, terete to bluntly subangled in cross-section, 1.1–1.7 × 0.9–1.2 mm, smooth, with smooth brown line from top to bottom on adaxial side.

Distribution

V12 17-distribution-map.jpg

Man., Colo., Ill., Ind., Iowa, Kans., La., Mich., Minn., Miss., Mo., Mont., N.Dak., N.Mex., Nebr., Okla., S.Dak., Tex., Utah, Wis., Wyo., n Mexico

Discussion

Varieties 2 (2 in the flora).

The two varieties of Euphorbia geyeri have been distinguished in large part by the presence of conspicuous involucral gland appendages in var. geyeri and the lack of appendages in var. wheeleriana. The two varieties are recognized here, but the variation in the size and presence of involucral gland appendages in the closely related E. bombensis suggests that this might be a somewhat variable character in this group of species.

Euphorbia geyeri is widespread throughout the central United States in sandy soils. Populations at the eastern edge of the range are often considered adventive (for example, sandy soils along railroad grades in Michigan). Euphorbia geyeri resembles E. glyptosperma (both being entirely glabrous), but that species has serrulate leaves (near the apex) and strongly angled, transverse-ridged seeds whereas E. geyeri has entire leaves and smooth, rounded seeds.

Selected References

None.

Key

1 Involucral gland appendages present; staminate flowers 5–9; seeds 1.1–1.4(–1.6) mm. Euphorbia geyeri var. geyeri
1 Involucral gland appendages absent or rudimentary; staminate flowers 10–20; seeds 1.6–1.7 mm. Euphorbia geyeri var. wheeleriana

"connate" is not a number. "distinct" is not a number."/2" is not declared as a valid unit of measurement for this property.

... more about "Euphorbia geyeri"
green +  and pale grayish +
emarginate +  and truncate +
0 cm0 mm <br />0 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
white to reddish-tinged +
pointed +  and rounded +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Victor W. Steinmann +, Jeffery J. Morawetz +, Paul E. Berry +, Jess A. Peirson +  and Ya Yang +
Engelmann +
pinnate +, symmetric +  and asymmetric +
rounded +, angled +  and asymmetric +
Anisophyllum +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
oblong;oblong-obovate or oblongelliptic +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
globose-ovoid +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.19 cm1.9 mm <br />0.0019 m <br />) +
Geyer’s sandmat +
in small , cymose clusters +  and solitary +
toothed +  and entire +
Man. +, Colo. +, Ill. +, Ind. +, Iowa +, Kans. +, La. +, Mich. +, Minn. +, Miss. +, Mo. +, Mont. +, N.Dak. +, N.Mex. +, Nebr. +, Okla. +, S.Dak. +, Tex. +, Utah +, Wis. +, Wyo. +  and n Mexico +
pistillate +  and staminate +
green +  and reddish +
0.2mm;0.4mm +
subtending +
elliptic-oblong +  and nearly circular +
0.2mm +  and 0.6mm +
axillary +  and terminal +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
campanulate +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
caducous +, deciduous +  and persistent +
bractlike +
top;bottom +
palmate +  and pinnate +
toothed +  and entire +
venation +  and conspicuous +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
connate;distinct +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Boston J. Nat. Hist. +
ashy white +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
terete +  and bluntly subangled +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
expanded +
free +, connate +  and distinct +
25 cm250 mm <br />0.25 m <br /> (45 cm450 mm <br />0.45 m <br />) +
mat-forming +
ascending +  and prostrate +
flattened +  and terete +
4 cm40 mm <br />0.04 m <br /> (25 cm250 mm <br />0.25 m <br />) +
connate +  and distinct +
interpetiolar +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
connate +  and distinct +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (?) +  and 0.06 cm0.6 mm <br />6.0e-4 m <br /> (?) +
Chamaesyce geyeri +
Euphorbia geyeri +
Euphorbia sect. Anisophyllum +
species +
axillary +  and terminal +
10 +, 9 +, 8 +, 7 +  and 6 +