Hypnum dieckei
Bot. Centralbl. 44: 423. 1890.
Plants medium-sized, dark green to golden green or nearly black. Stems 4–8 cm, redbrown to nearly black, procumbent to suberect, usually regularly pinnate, branches 0.2–1.5 cm; hyalodermis present, central strand poorly differentiated; pseudoparaphyllia foliose, broad. Stem-leaves falcate, curved downward toward substrate, ovatelanceolate, curving slightly to insertion, tapering to apex, 1–1.5 × 0.4–0.5 mm; base weakly decurrent; margins plane, entire to sinuate proximally, often sharply serrate toward apex; acumen narrow; costa usually indistinct; alar cells rectangular, abruptly enlarged, region well defined, 2 or 3 cells high, 3 or 4 cells wide, outermost cells with thinner walls; basal laminal cells shorter, wider than medial cells, yellowish to brownish, especially adjacent to hyaline alar cells, walls porose; medial cells 40–60 × 3–4 µm. Sexual condition dioicous; inner perichaetial leaves lanceolate, margins toothed, apex finely pointed, costa obscure. Seta redbrown when mature, 2–4.5 cm. Capsule nodding when mature, redbrown, cylindric, 3–4.5 mm; annulus 1-seriate or 2-seriate; operculum conic; endostome cilia rudimentary.
Phenology: Capsules mature Jun–Jul.
Habitat: Terrestrial, rock, logs, near water courses, seepage areas, open to somewhat shaded areas, usually where persistent moisture is available
Elevation: low to moderate elevations (0-1500 m)
Distribution
B.C., Alaska, Calif., Oreg., Wash., e Asia
Discussion
Hypnum dieckei is distributed around the north Pacific and can be frequent on logs or outcrops near streams. The plants are distinctive especially in summer when sporophytes are produced, because H. dieckei is the only North American species with nodding capsules. The dark stems and pinnate branching associated with excavate alar regions of thin-walled cells set off by inner thick-walled and pigmented basal cells are also usually enough to distinguish H. dieckei. Plants of H. dieckei have branches 0.5–1.5 mm wide. They are often firmly attached by rhizoids, but the rhizoids are usually absent from suberect plants; the alar cells are 40–65 × 20–30 µm and often bulging, with the outer cells somewhat collapsed inward; the supra-alar cells are few and triangular; and the basal laminal cells are rectangular.
Selected References
None.