familyOnagraceae
subfamilyOnagraceae subfam. Onagroideae
tribeOnagraceae tribe Onagreae
genusOenothera
sectionOenothera sect. Anogra
speciesOenothera deltoides
subspeciesOenothera deltoides subsp. howellii

Oenothera deltoides subsp. howellii

(Munz) W. M. Klein

Aliso 5: 180. 1962.

Warren L. Wagner
Basionym: Oenothera deltoides var. howellii Munz Aliso 2: 81. 1949
Treatment appears in FNA Volume 10.

Herbs perennial, densely strigillose, also villous and glandular puberulent distally; with relatively long, fleshy roots, grayish. Stems erect or ascending, several from base, not thickened near base, with numerous shorter lateral branches, these not encircling stems in older plants, 40–80 cm. Leaves cauline, without clear basal rosette, at least at anthesis; blade lanceolate, margins runcinate-pinnatifid. Flowers: buds fluted in distal 1/2, with free tips 1–9 mm; sepals 20–30 mm; petals 20–40 mm. Capsules 45–60 × 3–4 mm. 2n = 14.


Phenology: Flowering (Mar–)Jun–Jul(–Sep).
Habitat: Sand dunes and bluffs.
Elevation: 0–30 m.

Distribution

Calif.

Discussion

W. M. Klein (1964) determined subsp. howellii to be self-incompatible.

Subspecies howellii is known from the Antioch Dunes in Contra Costa County. Subspecies howellii is federally listed as endangered and is in the Center for Plant Conservation’s National Collection of Endangered Plants.

Selected References

None.

Lower Taxa

None.
... more about "Oenothera deltoides subsp. howellii"
RDF feed
2n chromosome quantity
14 +
4-locular septum architecture
incomplete +
4-locular septum architecture or structure in adjective form
1-locular +
Androecium position
epigynous +
Anther fixation
versatile +
Anther some measurement
0.5 cm5 mm <br />0.005 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Apex width
1.5mm +  and 5mm +
Author
Warren L. Wagner +
Authority
(Munz) W. M. Klein +
Basal leaf arrangement
alternate +
Basal leaf position
cauline +
Basal leaf presence
absent +
Basal rosette coloration
clear +
Basal rosette length
5 cm50 mm <br />0.05 m <br /> (25 cm250 mm <br />0.25 m <br />) +
Basal rosette prominence
conspicuous +
Basal rosette width
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Base coloration
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
Base shape
pedicel-like +
Basionyms
Oenothera deltoides var. howellii +
Berry architecture or shape
nutlike +
Berry dehiscence
indehiscent +  and loculicidal +
Blade margin shape
sinuate-dentate +  and pinnatifid denticulate subentire or entire +
Blade shape
lanceolate +
Branch atypical some measurement
40 cm400 mm <br />0.4 m <br /> (100 cm1,000 mm <br />1 m <br />) +
Branch orientation
decumbent +  and ascending +
Branch quantity
few +  and several +
Branch size
slender +
Branch some measurement
10 cm100 mm <br />0.1 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Bud orientation
nodding +
Bud shape
fluted +
Capsule architecture
sessile +
Capsule architecture or shape
nutlike +
Capsule architecture or structure in adjective form
4-locular +
Capsule arrangement or shape
contorted +
Capsule atypical length
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Capsule condition or texture
dry +
Capsule course
straight +  and curved +
Capsule dehiscence
dehiscent +
Capsule length
4.5 cm45 mm <br />0.045 m <br /> (6 cm60 mm <br />0.06 m <br />) +
Capsule orientation
spreading +
Capsule quantity
1/2 +
Capsule shape
tapering +, cylindrical +  and slightly 4-angled +
Capsule texture
woody +
Capsule width
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Carpel quantity
Many (?) +
Cauline rosette atypical length
12 cm120 mm <br />0.12 m <br /> (18 cm180 mm <br />0.18 m <br />) +
Cauline rosette length
4 cm40 mm <br />0.04 m <br /> (12 cm120 mm <br />0.12 m <br />) +
Cauline rosette width
0.5 cm5 mm <br />0.005 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Cel reproduction
vegetative +
Central stem architecture
unbranched +  and branched +
Central stem orientation
erect +
Central stem size or width
thickened +
Central stem texture
spongy +
Distribution
Calif. +
Elevation
0–30 m. +
Embryo-sac coating
oily +
Embryo-sac course
straight +
Embryo-sac presence
absent +
Embryo coating
oily +
Embryo course
straight +
Embryo presence
absent +
Endosperm coating
oily +
Endosperm course
straight +
Endosperm presence
absent +
Epidermis coloration
pink +  and white +
Epidermis relief
exfoliating +
Filament architecture
dithecal +
Filament fixation
basifixed +  and versatile +
Filament fusion
distinct +
Filament some measurement
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Floral-tube atypical some measurement
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Floral-tube duration
deciduous +
Floral-tube orientation
recurved +
Floral-tube presence
absent +
Floral-tube pubescence
hispid +, lanate +  and glabrous +
Floral-tube quantity
, +
Floral-tube some measurement
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flower architecture
zygomorphic +  and actinomorphic +
Flower architecture or arrangement or growth form
solitary +
Flower odor
unscented +
Flower part duration
deciduous +
Flower part life cycle
anthesis +
Flower quantity
1 +  and several +
Flower reproduction
bisexual +
Fruit architecture
winged +
Fruit course
curved +  and straight +
Fruit dehiscence
indehiscent +  and dehiscent +
Fruit shape
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Habitat
Sand dunes and bluffs. +
Hair height or length or size
short +
Hair position relational
interlocking +
Hair shape
curly +
Herb architecture
caulescent +
Herb architecture or function or pubescence
glandular +
Herb duration
perennial +
Herb growth form
suffrutescent +
Herb pubescence
puberulent +, villous +  and strigillose +
Indusium architecture
peltate +
Indusium prominence
conspicuous +
Inflorescence position
axillary +
Lateral branch height or length or size
short +
Lateral root orientation
spreading +
Leaf-axil orientation
nodding +  and erect +
Leaf architecture
simple +
Leaf arrangement
alternate +
Leaf atypical some measurement
13 cm130 mm <br />0.13 m <br /> (26 cm260 mm <br />0.26 m <br />) +
Leaf condition
later +
Leaf development
developed +
Leaf duration
deciduous +
Leaf position
cauline +
Leaf presence
absent +
Leaf some measurement
1 cm10 mm <br />0.01 m <br /> (13 cm130 mm <br />0.13 m <br />) +
Lobe arrangement or course or shape
linear +
Lobe quantity
4 (?) +
Locule arrangement or growth form
clustered +
Locule quantity
many +
Margin shape
runcinate-pinnatifid +
Mouth condition
closing +
Nectary presence
absent +
Number of lower taxa
0 +
Ovary architecture
pedicellate +  and sessile +
Ovary architecture or structure in adjective form
1-locular +
Ovary position
inferior +
Ovule architecture
bitegmic +
Ovule arrangement or growth form
clustered +
Ovule orientation
anatropous +
Ovule quantity
1 +  and 8 +
Perianth position
epigynous +
Petal arrangement
imbricate +
Petal arrangement or shape
convolute +
Petal coloration
fading pink +  and deep pink +
Petal presence
absent +
Petal quantity
4 +
Petal shape
obcordate +
Petal some measurement
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Petiole some measurement
1.5 cm15 mm <br />0.015 m <br /> (8 cm80 mm <br />0.08 m <br />) +
Phenology
Flowering (Mar–)Jun–Jul(–Sep). +
Plant life cycle
older +
Pollen-grain dehiscence or orientation
longitudinal +
Pollen arrangement
singly +
Polyad arrangement
singly +
Proximal part reproduction
sterile +
Publication title
Aliso +
Publication year
1962 +
Raphide quantity
abundant +
Reference
None +
Root coloration
grayish +
Root length or size
long +
Root position
lateral +
Root texture
fleshy +
Rosette development
developed +
Rosette position
basal +  and cauline +
Row atypical quantity
2 (4) +
Row quantity
1 +
Secretion condition
wet +
Secretion condition or texture
dry +
Secretion relief
papillate +
Secretion shape
clavate +  and globose +
Seed atypical quantity
1 (160) +
Seed coloration
buff +
Seed quantity
numerous +
Seed relief
sculptured +  and smooth +
Seed shape
obovoid +
Seed size
reduced +
Sepal arrangement or dehiscence
valvate +
Sepal atypical some measurement
1.3 cm13 mm <br />0.013 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Sepal coloration
not spotted +
Sepal duration
deciduous +
Sepal orientation
reflexed +
Sepal quantity
4 +
Sepal some measurement
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Septum architecture
incomplete +
Septum arrangement
parietal +, axile +  and placentation +
Septum fragility
fragile +
Septum presence
absent +
Septum width
thin +
Series length or size
longer +
Series quantity
2 +
Series size
unequal +
Series variability
equal +
Set length or size
longer +
Set quantity
2 +
Set variability
equal +
Shoot derivation
adventitious +
Shoot orientation
arising +
Shorter lateral branch quantity
numerous +
Shrub architecture
unarmed +
Shrub growth form
not clonal +
Shrub habitat
aquatic +, amphibious +  and terrestrial +
Slit dehiscence or orientation
longitudinal +
Spike architecture
leafy +
Stamen quantity
8 +
Stamen size
in 2 unequal series +  and subequal +
Stamen size or quantity
2 times as many or as many as sepals +
Stem architecture
with short , lateral branches +  and unbranched +
Stem growth form or orientation
decumbent +
Stem orientation
ascending +  and erect +
Stem quantity
several +
Stem size or width
not thickened +
Stem some measurement
40 cm400 mm <br />0.4 m <br /> (80 cm800 mm <br />0.8 m <br />) +
Stigma architecture
peltate +
Stigma development
developing +
Stigma quantity
1 +
Stigma shape
divided +
Stipule architecture
petiolate +  and sessile +
Stipule duration
caducous +
Stipule position
intrapetiolar +
Stipule presence
absent +
Stipule size
small +
Style pubescence
pubescent +  and glabrous +
Style quantity
1 +
Style some measurement
3.5 cm35 mm <br />0.035 m <br /> (6 cm60 mm <br />0.06 m <br />) +
Subshrub architecture
unarmed +
Subshrub growth form
not clonal +
Subshrub habitat
aquatic +, amphibious +  and terrestrial +
Surface architecture
smooth +, reticulate +  and furrowed +
Surface architecture or pubescence or relief
smooth +
Surface architecture or shape
entire +
Surface condition
wet +
Surface duration
persisting +
Surface life cycle
anthesis +
Surface prominence
obscured +
Surface quantity
7 (?) +
Surface relief
alveolate +
Surface shape
beaded +
Synonyms
Anogra deltoides +
Taxon family
Onagraceae +
Taxon name
Oenothera deltoides subsp. howellii +
Taxon parent
Oenothera deltoides +
Taxon rank
subspecies +
Tetrad arrangement
singly +
Tetrad dehiscence
3(-5)-aperturate +
Tip fusion
free +
Tip height or length or size
short +
Tip some measurement
0.1 cm1 mm <br />0.001 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Viscin thread dehiscence or orientation
longitudinal +
Viscin thread fusion
united +
Volume
Volume 10 +
Wall texture
woody +  and hard +
Wing architecture
smooth +  and erose +
Wing condition or texture
dry +
Wing relief
papillate +
Wing width
thick +