familyOnagraceae
subfamilyOnagraceae subfam. Onagroideae
tribeOnagraceae tribe Onagreae
genusOenothera
sectionOenothera sect. Oenothera
subsectionOenothera subsect. Candela
speciesOenothera heterophylla
subspeciesOenothera heterophylla subsp. heterophylla

Oenothera heterophylla subsp. heterophylla

Warren L. Wagner
Synonyms: Oenothera pyramidalis var. lindheimeri H. Léveillé
Treatment appears in FNA Volume 10.
Please click on the illustration for a higher resolution version.
Illustrator:

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Herbs densely to sparsely strigillose, also at least parts of inflorescence sparsely hirsute with spreading, pustulate-based hairs, and often glandular puberulent and villous. Flowers: buds with free tips spreading, 2–6 mm; floral-tube 25–42 mm; sepals 15–28 mm; petals 18–35 mm. 2n = 14.


Phenology: Flowering Jun–Sep.
Habitat: Sandy to sandy-loam soil of open sites in woodlands, with Persea borbonia, Pinus echinata, P. palustris, Quercus incana, Q. marilandica, Q. stellata, and Q. virginiana.
Elevation: 0–200 m.

Distribution

La., Tex.

Discussion

Populations of subsp. heterophylla were determined by W. Dietrich and W. L. Wagner (1988) to be self-incompatible. It occurs in a narrow range from eastern Texas (Austin, Bastrop, Brazos, Cass, Chambers, Cherokee, Dallas, Freestone, Gonzales, Gregg, Hardin, Harris, Henderson, Hopkins, Houston, Jasper, Lee, Leon, Liberty, Limestone, Nacogdoches, Newton, Robertson, Rusk, Sabine, San Augustine, San Jacinto, Smith, Sutton, Travis, Tyler, Upshur, Van Zandt, Victoria, Waller, and Wood counties) and south­western Louisiana (Caddo, Calcasieu, Erwin, Natchitoches,and Winn parishes). It is known from several historical specimens in St. Louis, Missouri, as an adventive but is apparently no longer growing in that area.

Oenothera variifolia Steudel is a superfluous name that pertains here.

Selected References

None.

Lower Taxa

None.
... more about "Oenothera heterophylla subsp. heterophylla"
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2n chromosome quantity
14 +
4-locular septum architecture
incomplete +
4-locular septum architecture or structure in adjective form
1-locular +
Androecium position
epigynous +
Anther fixation
versatile +
Anther some measurement
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Author
Warren L. Wagner +
Authority
Spach +
Basal leaf arrangement
alternate +
Basal leaf position
cauline +
Basal leaf presence
absent +
Basal stem length
7 cm70 mm <br />0.07 m <br /> (15 cm150 mm <br />0.15 m <br />) +
Basal stem width
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Base coloration
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
Base shape
pedicel-like +
Basionyms
Oenothera ser. +
Berry architecture or shape
nutlike +
Berry dehiscence
indehiscent +  and loculicidal +
Blade margin architecture or shape
serrate +
Blade margin shape
subentire +, dentate +, lobed to sinuate-dentate +  and lobed +
Blade shape
gradually narrowly lanceolate +  and lanceolate or elliptic +
Bract length or size
longer +
Bract some measurement
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Bud life cycle
mature +
Bud orientation
erect +
Bud shape
terete +  and weakly quadrangular +
Bud some measurement
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Capsule architecture
sessile +
Capsule architecture or shape
nutlike +
Capsule architecture or structure in adjective form
4-locular +
Capsule condition or texture
dry +
Capsule course
straight +  and curved +
Capsule course or shape
sigmoid +
Capsule dehiscence
indehiscent +  and dehiscent +
Capsule diameter
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Capsule length
1.3 cm13 mm <br />0.013 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Capsule shape
lanceoloid +
Capsule texture
woody +
Capsule width
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Carpel quantity
Many (?) +
Cauline stem length
3 cm30 mm <br />0.03 m <br /> (13 cm130 mm <br />0.13 m <br />) +
Cauline stem width
0.4 cm4 mm <br />0.004 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
Cel reproduction
vegetative +
Distribution
La. +  and Tex. +
Elevation
0–200 m. +
Embryo-sac coating
oily +
Embryo-sac course
straight +
Embryo-sac presence
absent +
Embryo coating
oily +
Embryo course
straight +
Embryo presence
absent +
Endosperm coating
oily +
Endosperm course
straight +
Endosperm presence
absent +
Epidermis coloration
whitish +  and green +
Epidermis relief
exfoliating +
Filament architecture
dithecal +
Filament fixation
basifixed +  and versatile +
Filament fusion
distinct +
Filament some measurement
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Floral-tube atypical some measurement
16.5 cm165 mm <br />0.165 m <br /> (19 cm190 mm <br />0.19 m <br />) +
Floral-tube course
straight +
Floral-tube duration
deciduous +
Floral-tube presence
absent +
Floral-tube pubescence
hispid +, lanate +  and glabrous +
Floral-tube quantity
, +
Floral-tube some measurement
2.5 cm25 mm <br />0.025 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
Flower architecture
zygomorphic +  and actinomorphic +
Flower architecture or arrangement or growth form
solitary +
Flower odor
unscented +
Flower part duration
deciduous +
Flower part life cycle
anthesis +
Flower quantity
2 +  and several +
Flower reproduction
bisexual +
Fruit architecture
winged +
Fruit course
curved +  and straight +
Fruit dehiscence
indehiscent +  and dehiscent +
Fruit shape
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Habitat
Sandy to sandy-loam soil of open sites in woodlands, with Persea borbonia, Pinus echinata, P. palustris, Quercus incana, Q. marilandica, Q. stellata, and Q. virginiana. +
Hair architecture
pustulate-based +
Hair height or length or size
short +
Hair orientation
spreading +
Hair position relational
interlocking +
Herb architecture
caulescent +
Herb duration
perennial +, short-lived +  and annual +
Herb growth form
suffrutescent +
Herb pubescence
strigillose +
Illustration
Present +
Indusium architecture
peltate +
Indusium prominence
conspicuous +
Inflorescence architecture or arrangement
lax +
Inflorescence density
dense +
Inflorescence orientation
erect +
Inflorescence position
axillary +
Inflorescence position or structure subtype
terminal +
Inflorescence pubescence
glabrate +, hirsute +, villous +, puberulent +  and glandular +
Lateral branch quantity
several +
Lateral root orientation
spreading +
Leaf-axil orientation
nodding +  and erect +
Leaf architecture
simple +
Leaf arrangement
alternate +
Leaf atypical some measurement
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Leaf condition
later +
Leaf duration
deciduous +
Leaf position
cauline +
Leaf presence
absent +
Leaf some measurement
3 cm30 mm <br />0.03 m <br /> (25 cm250 mm <br />0.25 m <br />) +
Lobe arrangement or course or shape
linear +
Lobe quantity
4 (?) +
Locule arrangement or growth form
clustered +
Locule quantity
many +
Margin shape
deeply lobed +  and remotely dentate or subentire +
Mouth condition
closing +
Nectary presence
absent +
Number of lower taxa
0 +
Ovary architecture
pedicellate +  and sessile +
Ovary architecture or structure in adjective form
1-locular +
Ovary position
inferior +
Ovule architecture
bitegmic +
Ovule arrangement or growth form
clustered +
Ovule orientation
anatropous +
Ovule quantity
1 +  and 8 +
Part architecture or function or pubescence
glandular +
Part pubescence
villous +, puberulent +  and hirsute +
Perianth position
epigynous +
Petal arrangement
imbricate +
Petal arrangement or shape
convolute +
Petal coloration
yellow +
Petal presence
absent +
Petal quantity
4 +
Petal shape
broadly elliptic +  and nearly rhombic +
Petal some measurement
1.8 cm18 mm <br />0.018 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
Phenology
Flowering Jun–Sep. +
Pollen-grain dehiscence or orientation
longitudinal +
Pollen arrangement
singly +
Pollen reproduction
fertile +
Polyad arrangement
singly +
Proximal part reproduction
sterile +
Publication title
Nouv. Ann. Mus. Hist. Nat. +
Publication year
1836 +
Raphide quantity
abundant +
Reference
None +
Rosette position
basal +  and cauline +
Row atypical quantity
2 (4) +
Row quantity
2 +
Secretion condition
wet +
Secretion condition or texture
dry +
Secretion relief
papillate +
Secretion shape
clavate +  and globose +
Seed atypical quantity
1 (160) +
Seed coloration
flecked +  and brown +
Seed length
0.11 cm1.1 mm <br />0.0011 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
Seed quantity
numerous +
Seed relief
sculptured +  and smooth +
Seed shape
ellipsoid +  and broadly ellipsoid +
Seed size
reduced +
Seed width
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Sepal architecture or dehiscence
splitting +
Sepal arrangement
separate +  and separating +
Sepal arrangement or dehiscence
valvate +
Sepal coloration
purple +, pink +, red +  and green +
Sepal duration
deciduous +
Sepal orientation
reflexed +
Sepal quantity
4 +
Sepal some measurement
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
Septum architecture
incomplete +
Septum arrangement
parietal +, axile +  and placentation +
Septum fragility
fragile +
Septum presence
absent +
Septum width
thin +
Series length or size
longer +
Series quantity
2 +
Series size
unequal +
Series variability
equal +
Set length or size
longer +
Set quantity
2 +
Set variability
equal +
Shoot derivation
adventitious +
Shoot development
developing +
Shoot orientation
arising +
Shrub architecture
unarmed +
Shrub growth form
not clonal +
Shrub habitat
aquatic +, amphibious +  and terrestrial +
Slit dehiscence or orientation
longitudinal +
Spike architecture
leafy +
Spike architecture or arrangement
lax +
Spike density
dense +
Stamen quantity
8 +
Stamen size
in 2 unequal series +  and subequal +
Stamen size or quantity
2 times as many or as many as sepals +
Stem architecture
branched +  and unbranched +
Stem growth form or orientation
decumbent +
Stem orientation
erect +
Stem some measurement
25 cm250 mm <br />0.25 m <br /> (70 cm700 mm <br />0.7 m <br />) +
Stigma architecture
peltate +
Stigma development
developing +
Stigma quantity
1 +
Stigma shape
divided +
Stipule architecture
petiolate +  and sessile +
Stipule duration
caducous +
Stipule position
intrapetiolar +
Stipule presence
absent +
Stipule size
small +
Style pubescence
pubescent +  and glabrous +
Style quantity
1 +
Style some measurement
4.5 cm45 mm <br />0.045 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
Subshrub architecture
unarmed +
Subshrub growth form
not clonal +
Subshrub habitat
aquatic +, amphibious +  and terrestrial +
Surface architecture
smooth +, reticulate +  and furrowed +
Surface architecture or coloration or relief
reticulate +
Surface architecture or shape
entire +
Surface condition
wet +
Surface duration
persisting +
Surface life cycle
anthesis +
Surface prominence
obscured +
Surface prominence or shape
flat +
Surface quantity
7 (?) +
Surface relief
pitted +
Surface shape
beaded +
Suture quantity
1 +
Synonyms
Oenothera pyramidalis var. lindheimeri +
Taxon family
Onagraceae +
Taxon name
Oenothera heterophylla subsp. heterophylla +
Taxon parent
Oenothera heterophylla +
Taxon rank
subspecies +
Tetrad arrangement
singly +
Tetrad dehiscence
3(-5)-aperturate +
Tip fusion
free +
Tip orientation
spreading +
Tip position or structure subtype
terminal +
Viscin thread dehiscence or orientation
longitudinal +
Viscin thread fusion
united +
Volume
Volume 10 +
Wall texture
woody +  and hard +
Wing architecture
smooth +  and erose +
Wing condition or texture
dry +
Wing relief
papillate +
Wing width
thick +