Silene uralensis subsp. porsildii

Bocquet

Candollea 22: 27. 1967.

Common names: Large-seeded nodding campion
Basionym: Melandrium macrospermum A. E. Porsild Rhodora 41: 225, plate 552, figs. 1–3. 1939
Synonyms: Gastrolychnis macrosperma (A. E. Porsild) Tolmatchew & Kozhanchikov Gastrolychnis soczaviana (Schischkin) Tolmatchew & Kozhanchikov Lychnis apetala var. macrosperma (A. E. Porsild) B. Boivin Lychnis macrosperma (A. E. Porsild) J. P. Anderson Melandrium soczavianum Schischkin Silene macrosperma (A. E. Porsild) Hultén Silene soczaviana (Schischkin) Bocquet
Treatment appears in FNA Volume 5. Treatment on page 211. Mentioned on page 168, 210.

Plants cespitose; taproot stout. Stems erect, simple or branched, stout, 10–35 cm, densely pubescent with long, purple-septate, eglandular and glandular-hairs. Leaves: basal numerous, blade 1.5–6 cm × 3–7 mm, somewhat fleshy, subglabrous; cauline in 1–3 pairs, blade narrowly lanceolate, 1.5–4 cm × 1–5 mm. Inflorescences sometimes forked, 1–3 (–4) -flowered. Pedicels erect or angled in flower and erect in fruit, stout, usually elongate, 1–10 cm, densely glandular-pubescent with long purple-septate hairs, ± viscid. Flowers erect; calyx veined, ovate-elliptic to broadly campanulate, 11–14 (–18) × 8–13 mm, veins brown or purple, densely glandular-pubescent with purple-septate hairs, lobes ovate-triangular, 2–3 mm, margins broad, membranous; corolla dingy purple-red, 11/3–11/2 times length of calyx, claw equaling calyx, limb obovate, 2-lobed, 2–4 mm. Capsules not contracted at mouth, equaling calyx; carpophore 1–2 mm. Seeds sooty brown, winged, round, flat, 2–2.5 mm diam.; wing inflated, broad, strongly rugose. 2n = 48.


Phenology: Flowering summer.
Habitat: Tundra, gravel slopes, talus, cliffs
Elevation: 0-2500 m

Distribution

V5 431-distribution-map.gif

Yukon, Alaska, Asia (Russian Far East), Asia (Siberia)

Discussion

The usually branched stout stems, erect flowers, broad capsule, and very large seeds with broad rugose wing distinguish subsp. porsildii.

Selected References

None.

Lower Taxa

None.

"/3-11/2timeslengthofcalyx" is not declared as a valid unit of measurement for this property.

eglandular +  and purple-septate +
awned +  and not hooded +
fimbriate;dissected;2-lobed;acute;obtuse +
John K. Morton +
Bocquet +
Melandrium macrospermum +
dentate +, emarginate to 2-fid +  and entire +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
subglabrous +
lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
reduced +
scarious +  and herbaceous +
1.4 cm14 mm <br />0.014 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
ovate-elliptic;broadly campanulate +
0.8 cm8 mm <br />0.008 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
slightly longer +
ovoid;globose +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
conspicuous +
Large-seeded nodding campion +
dingy purple-red +
dissected +  and variously--shaped +
dish--shaped +
expanded +
Yukon +, Alaska +, Asia (Russian Far East) +  and Asia (Siberia) +
0-2500 m +
peripheral +
subsessile +  and sessile +
conspicuous +
glandular-pubescent +
unisexual +  and bisexual +
stalked +  and simple +
eglandular +  and purple-septate +
Tundra, gravel slopes, talus, cliffs +
purple-septate +
sessile +  and petiolate +
whorled +  and opposite +
connate +
conspicuous +
2-lobed;obovate +
2mm +  and 4mm +
not differentiated +
purplish +  and green +
1 (?) +  and 4 (?) +
ovate-triangular +
pubescent;glabrous;ciliate +
membranous +
swollen +
3 +  and 5 +
crassinucellate +  and bitegmic +
campylotropous +
glandular-pubescent +
elongate +  and angled +
stout +  and slender +
1 cm10 mm <br />0.01 m <br /> (10 cm100 mm <br />0.1 m <br />) +
off-white tinged with purple +, dusky purple +, scarlet +, pink +  and white +
fugacious +
Flowering summer. +
sooty brown +
2mm;2.5mm +
papillate +  and tuberculate +
2.8 cm28 mm <br />0.028 m <br /> (4 cm40 mm <br />0.04 m <br />) +
persistent +
1 cm10 mm <br />0.01 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
Illustrated +
rhizomatous +  and stoloniferous +
fewer +  and 10 +
branched +  and simple +
prostrate +, decumbent +  and sprawling +
10 cm100 mm <br />0.1 m <br /> (35 cm350 mm <br />0.35 m <br />) +
[30 +, 4 +, 5 +  and 3 +
papillate +
distinct +
filiform +
0.15 cm1.5 mm <br />0.0015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Gastrolychnis macrosperma +, Gastrolychnis soczaviana +, Lychnis apetala var. macrosperma +, Lychnis macrosperma +, Melandrium soczavianum +, Silene macrosperma +  and Silene soczaviana +
Silene uralensis subsp. porsildii +
Silene uralensis +
subspecies +
divided +
10-30-veined +
purplish +, whitish +  and green +
inflated +, terete +, cylindric +  and campanulate urceolate or clavate +
membranous +
3 (?) +  and 5 (?) +
purple +  and brown +
glandular-pubescent +
stoloniferous +, rhizomatous +  and taprooted +
plant +  and cespitose +
pubescence of simple hairs or stalked glands +  and glabrous +
inflated +