Smelowskia porsildii

(W. H. Drury & Rollins) Jurtsev

Novosti Sist. Vyssh. Rast. 6: 309. 1970.

Basionym: Smelowskia calycina var. porsildii W. H. Drury & Rollins Rhodora 54: 105, fig. 3A. 1952
Synonyms: Hutchinsia calycina var. integrifolia Seemann Smelowskia calycina subsp. integrifolia (Seeman) Hultén Smelowskia calycina var. integrifolia (Seeman) Rollins Smelowskia jurtzevii Smelowskia spathulatifolia
Treatment appears in FNA Volume 7. Treatment on page 675. Mentioned on page 672, 673.

Plants sometimes canescent basally; caudex branched. Stems: several from base, unbranched, (0.3–) 0.5–1.4 (–2) dm, trichomes simple, 0.3–1 mm, mixed with smaller, dendritic ones, (sometimes simple ones absent). Basal leaves: petiole 0.7–3 (–4) cm, often ciliate, trichomes simple; blade usually linear to oblanceolate or spatulate, rarely obovate, (0.5–) 0.8–2.2 (–3.5) cm × 0.7–5 (–7) mm, margins usually entire, rarely apically 3 or 5-toothed or lobed, apex obtuse to rounded, (surfaces densely tomentose, grayish, trichomes mostly dendritic, mixed with fewer, simple ones, to 1 mm). Cauline leaves shortly petiolate; blade similar to basal, smaller distally, margins usually entire, rarely lobed. Racemes elongated in fruit. Fruiting pedicels divaricate to ascending, (forming 30–70˚ angle), proximalmost bracteate, 4–12 mm, pubescent, trichomes simple mixed with smaller, dendritic ones. Flowers: sepals 2.5–3.5 mm; petals white or creamy white, suborbicular to obovate, 4–6 × 2–3 (–4) mm, narrowed to claw, 1–2 (–3) mm, apex rounded; anthers oblong, 0.5–0.7 mm. Fruits divaricate to ascending, ellipsoid to oblong, subterete or slightly 4-angled, 6–10 × 1.5–2.5 mm, base and apex cuneate; valves each with prominent midvein; ovules 4–8 per ovary; style 0.1–0.5 mm. Seeds 1.5–2 × 0.9–1.2 mm. 2n = 12, 22, 24.


Phenology: Flowering Jun–Jul.
Habitat: Scree, dry gravelly slopes, sandstone shale-scree, fellfields, gravel beaches and benches, slides, alpine ridges, rock crevices and outcrops, sedge tundra, Dryas heath meadows, dry slopes, tundra slopes, conglomerate outcrops, marshes, sedge meadows, windswept sandstone ridges
Elevation: 0-1700 m

Distribution

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Alaska, e Asia (Russian Far East)

Discussion

As delimited here, Smelowskia porsildii is taken in the broad sense to include plants recognized by W. H. Drury Jr. and R. C. Rollins (1952) and Rollins (1993) as S. calycina var. integrifolia. The latter was raised by Velichkin to species rank, named as S. spathulatifolia because of the earlier-published S. integrifolia Bunge. Leaf shape and pedicel orientation, used by those authors to distinguish the two taxa, vary continuously. The species is also highly variable in the relative occurrence of simple versus dendritic trichomes and leaf margins. Although I. A. Al-Shehbaz and S. I. Warwick (2006) reduced S. spathulatifolia to synonymy of S. porsildii, the latter most likely represents a complex in which S. media might be involved. Conflicting chromosome numbers (e.g., 2n = 12, 18, 22, 24, 32) have been reported for this complex (Warwick and Al-Shehbaz 2006), and detailed cytological and molecular studies are needed to resolve the taxa involved. The complex involves diploid and tetraploid populations based on x = 6; the counts of 2n = 18 and 32 most likely reflect misidentifications.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Smelowskia porsildii"
24 +, 22 +  and 12 +
0.05 cm0.5 mm <br />5.0e-4 m <br /> - 0.07 cm0.7 mm <br />7.0e-4 m <br /> +
cuneate +, rounded +  and obtuse +
Ihsan A. Al-Shehbaz +
- W. H. Drury & Rollins Jurtsev +
cuneate +
Smelowskia calycina var. porsildii +
appendaged +  and unappendaged +
2.2 cm22 mm <br />0.022 m <br /> - 3.5 cm35 mm <br />0.035 m <br /> +
0.5 cm5 mm <br />0.005 m <br /> - 0.7 cm7 mm <br />0.007 m <br /> +
not auriculate +
0.8 cm8 mm <br />0.008 m <br /> - 2.2 cm22 mm <br />0.022 m <br /> +
entire +, 2-pinnatisect +  and 1-pinnatisect +
dissected +, repand +, sinuate +, crenate +  and dentate +
obovate;usually linear;oblanceolate or spatulate +
smaller +
0.07 cm0.7 mm <br />7.0e-4 m <br /> - 0.5 cm5 mm <br />0.005 m <br /> +
urceolate +, campanulate +  and tubular +
well-developed +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
accumbent +  and incumbent +
Alaska +  and e Asia - Russian Far East +
0-1700 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
stipitate +  and sessile +
0.6 cm6 mm <br />0.006 m <br /> - 1 cm10 mm <br />0.01 m <br /> +
ellipsoid +  and oblong subterete or slightly 4-angled +
0.15 cm1.5 mm <br />0.0015 m <br /> - 0.25 cm2.5 mm <br />0.0025 m <br /> +
Scree, dry gravelly slopes, sandstone shalScree, dry gravelly slopes, sandstone shale-scree, fellfields, gravel beaches and benches, slides, alpine ridges, rock crevices and outcrops, sedge tundra, Dryas heath meadows, dry slopes, tundra slopes, conglomerate outcrops, marshes, sedge meadows, windswept sandstone ridges sedge meadows, windswept sandstone ridges +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
lobed;lobed;5-toothed;entire +
smaller +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.2 cm2 mm <br />0.002 m <br /> - 0.3 cm3 mm <br />0.003 m <br /> +
0.3 cm3 mm <br />0.003 m <br /> - 0.4 cm4 mm <br />0.004 m <br /> +
creamy white +  and white +
rudimentary +
narrowed +, suborbicular +  and obovate +
0.1 cm1 mm <br />0.001 m <br /> - 0.2 cm2 mm <br />0.002 m <br /> +
0.2 cm2 mm <br />0.002 m <br /> - 0.3 cm3 mm <br />0.003 m <br /> +
3 cm30 mm <br />0.03 m <br /> - 4 cm40 mm <br />0.04 m <br /> +
0.7 cm7 mm <br />0.007 m <br /> - 3 cm30 mm <br />0.03 m <br /> +
Flowering Jun–Jul. +
trinucleate +  and 3 - -11-colpate +
Novosti Sist. Vyssh. Rast. +
suberect +, spreading +  and ascending +
slender +
mucilaginous +
aseriate +, biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.15 cm1.5 mm <br />0.0015 m <br /> - 0.2 cm2 mm <br />0.002 m <br /> +
oblong-lanceolate +  and oblong +
0.09 cm0.9 mm <br />9.0e-4 m <br /> - 0.12 cm1.2 mm <br />0.0012 m <br /> +
persistent +  and caducous +
distinct +
suberect +  and spreading +
0.25 cm2.5 mm <br />0.0025 m <br /> - 0.35 cm3.5 mm <br />0.0035 m <br /> +
perforated +  and complete +
reduced +
stipitate +  and sessile +
subterete +, terete +, angustiseptate +, 4-angled +, pyriform +, suboblong +, oblanceolate +, spatulate +, ellipsoid +, obovoid +, oblong +  and linear +
stipitate +  and sessile +
subterete +, terete +, angustiseptate +, 4-angled +, pyriform +, suboblong +, oblanceolate +, spatulate +, ellipsoid +, obovoid +, oblong +  and linear +
tetradynamous +
procumbent +  and decumbent +
erect +  and decumbent +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.01 cm0.1 mm <br />1.0e-4 m <br /> - 0.05 cm0.5 mm <br />5.0e-4 m <br /> +
Hutchinsia calycina var. integrifolia +, Smelowskia calycina subsp. integrifolia +, Smelowskia calycina var. integrifolia +, Smelowskia jurtzevii +  and Smelowskia spathulatifolia +
Smelowskia porsildii +
Smelowskia +
species +
forked +  and stellate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> - 0.1 cm1 mm <br />0.001 m <br /> +
coiled +, 1-7-veined +  and veined +
anastomosing +
0.4 cm4 mm <br />0.004 m <br /> - 1.2 cm12 mm <br />0.012 m <br /> +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +