Thelypodium howellii

S. Watson

Proc. Amer. Acad. Arts 21: 445. 1886.

EndemicConservation concern
Synonyms: Cartiera howellii (S. Watson) Greene Thelypodiopsis howellii (S. Watson) O. E. Schulz
Treatment appears in FNA Volume 7. Treatment on page 732. Mentioned on page 729.

Biennials or, rarely, perennials; (short-lived); glaucous, glabrous (except petioles). Stems branched distally, 1–8 (–9.3) dm. Basal leaves: petiole 0.5–2 cm, ciliate; blade often oblanceolate to spatulate, sometimes oblong or lanceolate, 2–10 (–13.5) cm × 10–23 (–40) mm, margins often lyrate to dentate, sometimes entire or repand. Cauline leaves (ascending); sessile; blade often lanceolate to linear, sometimes oblong, (0.8–) 1–9.7 (–14.5) cm × 2–10 (–30) mm, (base usually amplexicaul to strongly sagittate, rarely auriculate), margins entire. Racemes lax, elongated in fruit, (flower buds ovate to lanceolate). Fruiting pedicels usually ascending, rarely divaricate-ascending, usually straight, rarely slightly incurved, stout, (2.5–) 3.5–8 (–14.5) mm, not flattened at base. Flowers: sepals erect, linear or lanceolate, (3.5–) 5–8.5 (–11.5) × (0.7–) 1–2 (–2.5) mm; petals lavender to purple, linear or oblanceolate to spatulate, (6–) 9–16.5 (–22) × 0.5–2.7 (–3) mm, margins partially crisped, claw differentiated from blade, [slender, (2.5–) 4–7 (–11) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments slightly tetradynamous, median pairs (3–) 5.5–9 (–12) mm, lateral pair (2.5–) 4.5–8 (–10) mm; anthers slightly exserted to included, linear, (1.5–) 2–4 (–5.5) mm, not circinately coiled; gynophore (stout), 0.5–1 (–3.5) mm. Fruits often ascending, sometimes erect or divaricate, slightly torulose, straight or slightly incurved (forming straight line with pedicel), terete, 1.5–6.5 (–8.2) cm × 1–1.3 (–2) mm; ovules 22–40 per ovary; style cylindrical or subclavate, (0.5–) 1–2.7 (–4.3) mm. Seeds (1–) 1.2–1.7 (–2) × 0.5–0.7 mm.

Discussion

Subspecies 2 (2 in the flora).

Streptanthus howellii (S. Watson) M. E. Jones is a later homonym of S. howellii S. Watson 1885.

Selected References

None.

Key

1 Petals usually linear to narrowly oblanceolate, rarely spatulate, 0.5-1.2(-3) mm wide; median filaments connate most of their lengths; ne California, c, s Oregon, s Washington. Thelypodium howellii subsp. howellii
1 Petals usually spatulate, rarely oblanceolate, (1.2-) 1.5-2.7(-3) mm wide; median filaments distinct; e Oregon. Thelypodium howellii subsp. spectabilis

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Thelypodium howellii"
0.4 cm4 mm <br />0.004 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
slightly exserted +  and included +
usually linear +  and linear-oblong rarely oblong or ovate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Ihsan A. Al-Shehbaz +
S. Watson +
not +  and rosulate +
straight +
incurved +, divaricate-ascending +  and ascending +
not flattened +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Pachypodium +
appendaged +  and unappendaged +
9.7 cm97 mm <br />0.097 m <br /> (14.5 cm145 mm <br />0.145 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
not +  and auriculate +
1 cm10 mm <br />0.01 m <br /> (9.7 cm97 mm <br />0.097 m <br />) +
laciniate +, lobed +, lyrate +  and dentate +
oblong;lanceolate;linear +
much smaller +  and reduced +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
accumbent +  and incumbent +
Calif. +, Oreg. +  and Wash. +
straight +  and curved +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
connate +  and distinct +
lateral +  and median +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
zygomorphic +  and actinomorphic +
0.1 cm1 mm <br />0.001 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
purple;lavender;purple;lavender;white +
dilated +, spatulate +  and obovate or oblanceolate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
6.5 cm65 mm <br />0.065 m <br /> (8.2 cm82 mm <br />0.082 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
straight +
1.5 cm15 mm <br />0.015 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
incurved +, erect +  and ascending +
terete +  and torulose +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
crisped;repand;entire;lyrate to dentate;lyrate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
1.65 cm16.5 mm <br />0.0165 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
lavender +  and purple +
9mm;16.5mm +
rudimentary +
oblanceolate +  and spatulate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
trinucleate +  and 3(-11)-colpate +
Proc. Amer. Acad. Arts +
reflexed +, erect +  and ascending +
stout +  and slender +
mucilaginous +
aseriate +, uniseriate +  and biseriate +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
ovate +, oblong +  and flattened +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
not basally +  and saccate +
0.85 cm8.5 mm <br />0.0085 m <br /> (1.15 cm11.5 mm <br />0.0115 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
persistent +  and caducous +
distinct +
lanceolate +  and linear +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
Endemic +  and Conservation concern +
tetradynamous +
subequal +
procumbent +  and decumbent +
entire +  and capitate +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.43 cm4.3 mm <br />0.0043 m <br />) +
persistent +
distinct +
subclavate +  and cylindrical +
0.1 cm1 mm <br />0.001 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
Cartiera howellii +  and Thelypodiopsis howellii +
Thelypodium howellii +
Thelypodium +
species +
dendritic +  and simple +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +  and biennial +
glaucous +  and glabrous +
perennial +, biennial +  and annual +
subshrub +  and shrub +
aquatics +  and terrestrial +
glabrous +  and pubescent +