Codriophorus sect. Codriophorus

Synonyms: Racomitrium sect. Papillosa Chevallier Trichostomum sect. Stenotrichum
Treatment appears in FNA Volume 27. Treatment on page 296.
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Stem sparingly or freely irregularly, dichotomously or fasciculately branched, sometimes filiform and almost unbranched, absent short, lateral tuft-like branchlets. Leaves lingulate, elliptic, broadly ovate, ovate or oblong-lanceolate to broadly or occasionally narrowly lanceolate; margins 1–2 (–4) -stratose, recurved, reflexed, incurved or plane on both sides; apices narrowly to broadly rounded-obtuse or long acuminate to acute, muticous or very rarely shortly hyaline-tipped, entire or sharply to bluntly, remotely irregularly toothed, erose-dentate, sinuate to cristate in the upper 1/5–1/4; costa strong and broad, ceasing in mid leaf to subpercurrent, in transverse-section 2–4-stratose in the distal and medial parts, weakly to strongly convex, reniform, crescent-shaped to semiterete abaxially, flat or convex on the adaxial side, in the basal part 3–6 (–7) -layered, situated at the bottom of a shallow or deeper, usually wide-angled groove, lunate, reniform or rectangular and distinctly flattened, strongly convex on the abaxial side; basal marginal laminal cells not differentiated; distal laminal cells 1-stratose to variously 2-stratose, irregularly roundedquadrate, subquadrate, oval to rounded-hexagonal, strongly papillose to smooth, straight-walled in the distal part.

Distribution

North America, South America (Colombia), Eurasia, n Africa, Atlantic Islands (Azores), Atlantic Islands (Canary Islands), Atlantic Islands (Iceland), Atlantic Islands (Madeira)

Discussion

Species 9 (6 in the flora).

Members of sect. Codriophorus are recognized by their lingulate, broadly ovate to ovate- or oblong-lanceolate leaves, isodiametric and straight-walled distal laminal cells, and costa that is broad and robust, usually over 100 µm wide at the base. The costa is situated in a shallow, wide-angled groove and is mostly 2–4-stratose in the distal and medial parts, 3–6(–7)-stratose in the basal part, and anatomically strongly differentiated.

Selected References

None.

Key

1 Laminal cells smooth or very slightly papillose on young leaves; leaf margins entire or with a few irregular teeth at the apex. Codriophorus depressus
1 Laminal cells distinctly papillose; leaf margins remotely irregularly toothed, erose-dentate, sinuate to cristate in the distal 1/5-1/4, if entire then broadly rounded-obtuse > 2
2 Leaves lanceolate; apex irregularly bluntly erose-dentate; costa markedly convex and crescent-shaped abaxially. Codriophorus ryszardii
2 Leaves lingulate, elliptical to broadly ovate or ovate-lanceolate; apex broadly rounded-obtuse to acute, irregularly dentate to entire > 3
3 Costa subpercurrent, entire and only shortly forked at the tip > 4
3 Costa extending 1/2-3/4 way up the leaf, distinctly spurred > 5
4 Laminal cells strongly papillose, 1- or very rarely 2-stratose; leaves unbordered. Codriophorus acicularis
4 Laminal cells indistinctly papillose over the cell walls, always variously 2-stratose in the distal part; leaves distinctly bordered with fleshy and broad limbidia. Codriophorus norrisii
5 Leaf apex broadly rounded, flat to subcucullate, always entire or sometimes sinuate to remotely bluntly erose-dentate. Codriophorus mollis
5 Leaf apex broadly rounded-obtuse to subacute, often tubular, strongly dentate or very rarely subentire. Codriophorus aduncoides
hyaline-tipped +
entire or +  and sharply bluntly +
Halina Bednarek-Ochyra +  and Ryszard Ochyra +
P. Beauvois +
both +  and papillose +
toothed +
short to long +
nodulose;oblate;quadrate;elongate +
thick-walled +  and nodulose-porose +
rectangular;linear +
3-6(-7)-layered +
divided +
papillose +
undifferentiated +
naked +, plicate +  and smooth +
0 +  and 1/2 +
distinctly verrucose +  and papillose +
conic-mitrate +
small;large +
sulcate +  and smooth +
nearly +  and symmetric +
yellowbrown +, redbrown +  and brown +
striate +  and smooth +
ovoid;long-cylindric +
2-5-stratose;percurrent +
spurred +  and entire +
subpercurrent +
papillose +  and smooth +
convex +, flattened +, rectangular +, reniform +, lunate +, crescent-shaped +  and semiterete abaxially +
much larger +
straight-walled +, 1-stratose +  and variously 2-stratose +
papillose to smooth +  and papillose +
North America +, South America (Colombia) +, Eurasia +, n Africa +, Atlantic Islands (Azores) +, Atlantic Islands (Canary Islands) +, Atlantic Islands (Iceland) +  and Atlantic Islands (Madeira) +
sinuose-nodulose +
isodiametric +  and elongate +
wide-angled +
elliptical +  and lingulate +
porose +, 2-stratose +  and 1-stratose +
elliptical +  and oblong-elliptical +
thick-walled +  and sinuose +
dioicous +  and sexual +
curved +  and straight +
recurved +, spreading +  and erect +
oblong-lanceolate +  and broadly or occasionally narrowly lanceolate +
1-2(-4)-stratose +
erose-dentate to papillose-crenulate or crenate +  and entire +
incurved +, reflexed +  and recurved +
subpercurrent +
long-rostrate +
lanceolate;linear +
Mém. Soc. Linn. Paris +
straight +
erect +, sinistrorse +  and dextrorse +
one;several +
papillose +  and smooth +
short to long +
ornamented +  and smooth +
spheric +
unbranched +  and branched +
creeping +  and ascending +
filiform +
Racomitrium sect. Papillosa +  and Trichostomum sect. Stenotrichum +
Codriophorus sect. Codriophorus +
Codriophorus +
section +
nodulose +
incrassate +
usually olivaceous;blackish green +
robust +  and gracile +