Crataegus intricata

Lange

Bot. Tidsskr. 19: 264. 1895.

Common names: Intricate hawthorn
Endemic
Synonyms: Crataegus boyntonii Beadle C. foetida Ashe C. intricata var. boyntonii (Beadle) Kruschke
Treatment appears in FNA Volume 9. Treatment on page 588. Mentioned on page 586, 587, 589, 591.

Shrubs, 10–60 dm. Stems: twigs: new growth reddish, glabrous, 1–2-years old dark reddish black, older grayer; thorns on twigs absent or frequent, straight to recurved, 2-years old blackish, slender, 2.5–5 cm. Leaves: petiole 1 mm wide, length 40–50% blade, usually glabrous, sometimes hairy, glandular; blade broadly elliptic to broadly ovate or oblong, 4–8 (–9) cm, base broadly cuneate to ± rounded, lobes 3–5 per side, sinuses ± shallow to moderately deep, angled to rounded, lobe apex acute to acuminate, often reflexed, margins ± serrate, teeth gland-tipped at least young, veins 4–6 per side, apex acute, surfaces glabrous. Inflorescences 3–8-flowered; branches glabrous; bracteoles membranous to nearly herbaceous, margins short-stipitate-glandular. Flowers 16–20 mm diam.; hypanthium glabrous; sepals 6 mm, margins glandular-serrate; stamens 10, anthers ivory or cream; styles 3–5. Pomes dull yellow to orange or russet, sometimes green mature, orbicular to ± pyriform, 8–13 mm diam., glabrous; sepals on collar, patent-reflexed; pyrenes 3–5.2n = 51.


Phenology: Flowering late Apr, early Jun; fruiting Sep–Oct.
Habitat: Open forests, forest gaps, open places
Elevation: 10–300 m

Distribution

V9 1001-distribution-map.jpg

Ont., Ala., Conn., Del., Ga., Ill., Ind., Ky., Md., Mass., Mich., Mo., N.J., N.Y., N.C., Ohio, Pa., R.I., S.C., Tenn., Vt., Va., W.Va.

Discussion

Crataegus intricata ranges in Appalachia from southern New England to Georgia and Alabama. The distribution given above, relative to similar taxa, may be somewhat over-represented because the species has been the default identification for doubtful (particularly fruiting) material of much ser. Intricatae with glabrous leaves and inflorescences.

Crataegus intricata is the common, widespread member of this series with lobed leaves, ten stamens, white or cream anthers, and glabrous inflorescences. Depth and sharpness of lobes and location of widest part of leaf are somewhat variable, as is plant stature, some forms appearing mature at 1–2 m.

W. W. Eggleston collected a distinctive form in the Crataegus intricata complex in September 1908 from North Carolina, in which the short-shoot leaves are broadly ovate to deltate with shallow obtuse lobes. This has a printed label naming it C. virgata Ashe. Though type material of C. virgata could not be located, the protologue by Ashe could fit. It is possible that all the southeastern forms from the intricata complex with shallow and obtuse lobes, proportionately wide leaves, and cream anthers represent C. virgata.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."glabrous" is not a number."adnate" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Crataegus intricata"
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
cream +  and ivory +
James B. Phipps +
orangebrown +
thin-exfoliating +  and corrugated +
compound +  and simple +
opposite +  and alternate +
broadly cuneate +  and more or less rounded +
Intricatae +
8 cm80 mm <br />0.08 m <br /> (9 cm90 mm <br />0.09 m <br />) +
broadly elliptic +  and broadly ovate or oblong +
4 cm40 mm <br />0.04 m <br /> (8 cm80 mm <br />0.08 m <br />) +
coriaceous +
membranous;nearly herbaceous +
adnate +  and distinct +
crowded +, scattered +  and alternate +
Intricate hawthorn +
Ont. +, Ala. +, Conn. +, Del. +, Ga. +, Ill. +, Ind. +, Ky. +, Md. +, Mass. +, Mich. +, Mo. +, N.J. +, N.Y. +, N.C. +, Ohio +, Pa. +, R.I. +, S.C. +, Tenn. +, Vt. +, Va. +  and W.Va. +
not +  and aggregated +
10–300 m +
less hard +  and mealy +
1.6 cm16 mm <br />0.016 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
not +  and aggregated +
yellow +  and red or purplish +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
black +, dark reddish +  and reddish +
older +, old +  and new +
Open forests, forest gaps, open places +
constricted +
crassinucellate +
deciduous +
craspedodromous +
glandular-serrate +  and serrate +
short-stipitate-glandular +
glandular-serrate +  and glandular-pectinate +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
elliptic +  and circular +
0 (?) +  and 4 (?) +
pale paper brown +
free +  and distinct +
post-mature +
stipitate +  and sessile +
1 cm10 mm <br />0.01 m <br /> (?) +
Flowering late Apr, early Jun +  and fruiting Sep–Oct. +
adnate +, free +, connate +  and distinct +
dark gray-brown +  and russet +
exfoliating +  and corrugated +
pruinose +
green +, yellow +  and orange or russet +
0.8 cm8 mm <br />0.008 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
orbicular +  and more or less pyriform +
Bot. Tidsskr. +
not arillate +
persistent +
free +  and distinct +
patent-reflexed +
triangular +
nonaccrescent +
6 cm60 mm <br />0.06 m <br /> (?) +
1-50-flowered +  and few-leaved +
less shallow +  and moderately deep +
free +  and distinct +
branched +  and simple +
persistent +
distinct +
exsert +  and lateral +
elongate +
Crataegus boyntonii +, C. foetida +  and C. intricata var. boyntonii +
Crataegus intricata +
Crataegus (sect. Coccineae) ser. Intricatae +
species +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
straight +
slender +
2.5 cm25 mm <br />0.025 m <br /> (5 cm50 mm <br />0.05 m <br />) +
gland-tipped +
inconspicuous +
buff to gray-brown +  and corrugated +
less erect +  and oblique +
1 +  and several +
flexuous +  and straight +
determinate +
4 (?) +  and 6 (?) +