Eleocharis uniglumis

(Link) Schultes

Mant. 2: 88. 1824.

Common names: Éléocharide unigume
Illustrated
Basionym: Scirpus uniglumis Link Jahrb. Gewächsk. 1(3): 77. 1820
Synonyms: Eleocharis halophila (Fernald & Brackett) Fernald Eleocharis uniglumis var. halophila Fernald & Brackett
Treatment appears in FNA Volume 23. Treatment on page 76. Mentioned on page 65, 72, 73, 75, 78.

Plants perennial, mat-forming; rhizomes evident, long, 0.3–1 mm thick, soft to firm, cortex often fugaceous, longer internodes 10–25 mm, scales fugaceous, 5–6 mm, membranous, not fibrous. Culms terete, often with some blunt ridges when dry, (5–) 10–60 cm × 0.2–1.5 mm, firm, internally spongy. Leaves: distal leaf-sheaths persistent, not splitting, proximally red, distally stramineous to green, often callose, thinly papery to thickly membranous, apex often dark redbrown, obtuse to subacute, tooth absent. Spikelets ovoid to lanceoloid, 5–10 × 2–3 (–4) mm, apex acute; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales often spreading in fruit, 10–20, 3–4 per mm of rachilla, brown to often redbrown, midrib regions mostly stramineous to green, broadly ovate, 3–4 × 1.8–2.5 mm, entire, apex acute to obtuse, often some carinate in distal part of spikelet. Flowers: perianth bristles 0–4 (–5), light-brown to stramineous, stout, usually unequal, rudimentary to equaling achene; stamens 3; anthers dark yellow to stramineous, 1.2–2 mm; styles 2-fid. Achenes not persistent, dark yellow or medium or dark-brown, ellipsoid, obovoid, or obpyriform, biconvex, angles obscure, 1.3–1.8 × 1–1.4 mm, apex rounded, neck absent or short, smooth at 30X, or sometimes finely rugulose at 10–20X with 20 or more horizontal ridges in vertical series. Tubercles brown to whitish, pyramidal, much higher than wide to slightly depressed, sometimes spongy and with vertical rows of depressions, 0.4–0.8 × 0.3–0.8 mm.


Phenology: Fruiting summer.
Habitat: Mostly coastal, brackish (to fresh?) shores, marshes
Elevation: 0–2300 m

Distribution

V23 101-distribution-map.jpg

Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Ont., P.E.I., Que., Colo., Del., Maine, Mass., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.C., N.Dak., R.I., S.Dak., Utah, Va., Eurasia

Discussion

Plants treated as Eleocharis uniglumis fall within the large morphologic variation of Eurasian E. uniglumis. Two subspecies and 3 varieties were recognized for northern Europe (S.-O. Strandhede 1966), and five species that were recognized by I. D. Zinserling (1935) were placed in synonymy under E. uniglumis (S.-O. Strandhede 1966). Recognition of infraspecific taxa within North American E. uniglumis is premature. Plants commonly called E. halophila or E. uniglumis var. halophila, found mostly in Atlantic Coastal brackish habitats, have floral scales that are usually narrower and more densely placed on the rachilla than plants usually called E. uniglumis, which are found mostly in the interior; some plants are intermediate in expression of these characters. The achene and tubercle shape characters used by M. L. Fernald (1950) to distinguish E. uniglumis from E. halophila are not valid. In North America, E. uniglumis is difficult to separate from E. erythropoda and E. kamtschatica, in both of which the spikelets have only the proximal scale without a flower (empty) and the proximal scale completely amplexicaulous. Eleocharis uniglumis differs from E. erythropoda only in its broader floral scales, which are less densely placed on the rachilla; it differs from E. kamtschatica only in its smaller tubercles. It is also difficult to separate from some specimens of E. macrostachya in which the spikelets have proximal scales that are sometimes completely amplexicaulous; such plants differ from E. uniglumis only in the absence of a flower in the axil of the subproximal scale of some of the spikelets. I have not seen voucher specimens for the chromosome numbers of 2n = 27 and 28 reported by S.-O. Strandhede (1967) from Massachusetts and Nebraska, which are lower than the 2n = (44–)46(47–88) reported for Europe (S.-O. Strandhede 1965).

Selected References

None.

Lower Taxa

None.

"shortened" is not a number.

... more about "Eleocharis uniglumis"
dark-brown +  and dark yellow +
not persistent +
cancellate or reticulate +  and rugulose +
biconvex;obpyriform;obovoid;obpyriform;obovoid;ellipsoid +
color +  and texture +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
1mm;1.4mm +
dark yellow;stramineous +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
dark redbrown +
rounded;carinate;acute;obtuse +
S. Galen Smith* +, Jeremy J. Bruhl* +, M. Socorro González-Elizondo* +  and Francis J. Menapace* +
(Link) Schultes +
flattened +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
persistent +
enlarged +
Scirpus uniglumis +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
ascending +  and appressed +
parallel +  and divergent +
terete +, rolled +  and plicate +
Éléocharide unigume +
0 cm0 mm <br />0 m <br /> (100 cm1,000 mm <br />1 m <br />) +
0.3mm;5mm +
distally stramineous +  and green +
Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Ont. +, P.E.I. +, Que. +, Colo. +, Del. +, Maine +, Mass. +, Nebr. +, Nev. +, N.H. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, N.Dak. +, R.I. +, S.Dak. +, Utah +, Va. +  and Eurasia +
0–2300 m +
redbrown +  and brown to often +
membranous +  and thinly papery +
hypogynous +  and subtending +
biconvex +  and trigonous +
Mostly coastal, brackish (to fresh?) shores, marshes +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
flattened +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
rugulose +
with (1-)3-6(-30) bristles and/or scales +
Fruiting summer. +
2-3(-4)-carpellate +
3 (?) +  and 1 (?) +
stramineous;medium brown or red brown or blackish brown +
ascending +  and horizontal +
caudex-like +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
10cm;60cm +
spongy;firm +
0.2mm;1.5mm +
adventitious +
fugaceous +
basal +  and proximal +
2-keeled +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
not fibrous +  and membranous +
cylindric +
Illustrated +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
ovoid;lanceoloid +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
Eleocharis halophila +  and Eleocharis uniglumis var. halophila +
Eleocharis uniglumis +
Eleocharis (sect. Eleocharis) ser. Eleocharis +
species +
brown;whitish +
0.4mm;0.8mm +
much higher than wide to slightly depressed +
pyramidal +
0.3mm +  and 0.8mm +
plant +  and mat-forming +