Orobanche californica subsp. jepsonii

(Munz) Heckard

Madroño 22: 57. 1973.

Endemic
Basionym: Orobanche grayana var. jepsonii Munz Bull. Torrey Bot. Club 57: 617, plate 38, fig. 10. 1931
Synonyms: Aphyllon californicum subsp. jepsonii (Munz) A. C. Schneider
Treatment appears in FNA Volume 17. Treatment on page 480. Mentioned on page 478, 481.

Plants simple or branched proximally, (8–) 10–35 cm, portion proximal to inflorescence 6–15 cm, slender. Inflorescences racemes, sometimes subcorymbose, 5–20 cm; bracts pallid to pinkish tinged, drying brown. Pedicels 3–20 mm. Calyces (10–) 15–24 (–27) mm, lobes pallid or pinkish tinged, subulate to linear-subulate, (8–) 10–20 (–25) mm. Corollas whitish to pinkish distally, often with veins of lips deep rose or purplish, (25–) 30–35 (–40) mm; tube slender, abruptly widening toward throat; throat 8–10 mm wide at base of lobes; lips 10–12 (–15) mm, abaxial lobes usually narrowly lanceolate, 3–5 mm wide, apex acute, adaxial lobes deltate, apex acute, often erosulate or emarginate. 2n = 48.


Phenology: Flowering May–Oct.
Habitat: Bottomlands, dry hillsides.
Elevation: 100–2000 m.

Discussion

Subspecies jepsonii is distributed primarily in the Central Valley and surrounding mountains from the Cascade Range of Plumas and Shasta counties in the Sierra Nevada to Kern County and the central California Coast Ranges from Santa Cruz to Tehama counties.

The most frequently reported hosts for subsp. jepsonii are Baccharis, Grindelia, and Solidago (Asteraceae). Reports of Eriophyllum are unconfirmed, and reports of non-Asteraceae hosts such as Eriogonum and Rubus are highly unlikely.

Subspecies jepsonii is circumscribed similarly to what Munz described as Orobanche grayana var. jepsonii. It includes foothill and montane plants that resemble subsp. grayana in size and corymbose inflorescence, albeit with a deeper colored corolla and a preference for dry, rocky habitats.

L. R. Heckard (1973) indicated that subsp. jepsonii and subsp. grayana are very similar both geographically and morphologically. The two subspecies are separated by minor differences in plant size, corolla color and shape, and habitat.

Selected References

None.

Lower Taxa

None.
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
glabrous +  and villous +
emarginate +  and erosulate +
glandular-pubescent +
L. Turner Collins +, Alison E. L. Colwell +  and George Yatskievych +
(Munz) Heckard +
enlarged +
Orobanche grayana var. jepsonii +
oblong;lanceolate;oblong;lanceolate;deltate;triangular;ovate +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
not leathery +, not +  and fleshy +
brown;pallid;pinkish tinged +
appressed +  and spreading +
less lanceolate +  and oblanceolate +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2.4 cm24 mm <br />0.024 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
pallid +  and dark purple pink yellow or white +
divided +
1.5 cm15 mm <br />0.015 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
ovoid;cylindric-ovoid +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
3.5 cm35 mm <br />0.035 m <br /> (4 cm40 mm <br />0.04 m <br />) +
purplish +, rose +, whitish +  and pinkish +
bent +  and tubular +
constricted +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
spikelike +
Calif. +  and Oreg. +
100–2000 m. +
pubescent +  and glabrous +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Bottomlands, dry hillsides. +
axillary +  and terminal +
6 cm60 mm <br />0.06 m <br /> (15 cm150 mm <br />0.15 m <br />) +
branched +  and simple +
pallid cream +  and nearly white +
5 cm50 mm <br />0.05 m <br /> (20 cm200 mm <br />0.2 m <br />) +
deciduous +
1.2 cm12 mm <br />0.012 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
cream +  and pinkish or purplish tinged or pink +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
pinkish tinged +  and pallid +
loosely ascending +  and recurved +
glandular-pubescent +
subulate +  and linear-subulate +
erosulate +  and entire +
lobed +, toothed +  and entire +
tenuinucellate +  and unitegmic +
campylotropous-like +
spikelike +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering May–Oct. +
subcorymbose +
spikelike +
branched +  and unbranched +
inconspicuous +
coralloid +
stout +  and slender +
black;tan;dark-brown +
500 +  and 2000 +
prismatic +, ovoid +  and oblong-ellipsoid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
5 +  and 4 +
subequal +
yellow +  and white +
aerial +  and subterranean +
puberulent +  and glabrous +
capitate +  and crateriform-peltate +
clavate +  and 2-4-lobed +
Aphyllon californicum subsp. jepsonii +
Orobanche californica subsp. jepsonii +
Orobanche californica +
subspecies +
8mm;10mm +
cream;pinkish or purplish tinged or pink +
glandular-puberulent +
slender +
widening +
branched +  and simple +
8 cm80 mm <br />0.08 m <br /> (10 cm100 mm <br />0.1 m <br />) +
perennial +  and annual +
holoparasitic +  and hemiparasitic +
stout +  and slender +