Penstemon heterodoxus

A. Gray in A. Gray et al.

Syn. Fl. N. Amer. 2(1): 269. 1878.

Common names: Sierra beardtongue
Endemic
Treatment appears in FNA Volume 17. Treatment on page 207. Mentioned on page 189, 208.
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Herbs. Stems decumbent, ascending, or erect, 5–65 cm, glabrous or retrorsely hairy, sometimes also glandular-pubescent distally, not glaucous. Leaves basal and cauline, not leathery, glabrous; basal and proximal cauline 5–140 × 3–20 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–5 pairs, short-petiolate or sessile, 10–60 (–70) × 1–14 mm, blade oblong to oblanceolate or lanceolate, base tapered to clasping, margins entire, apex obtuse to acute. Thyrses interrupted, cylindric, 1–12 cm, axis ± glandular-pubescent, verticillasters 1–6, cymes (1 or) 2–6-flowered, 2 per node; proximal bracts lanceolate, 4–35 × 1–6 mm, margins entire; peduncles and pedicels erect, ± glandular-pubescent. Flowers: calyx lobes oblong to lanceolate, 4–6 × 0.7–1.2 mm, glandular-pubescent; corolla lavender to blue, violet, or purple, essentially without nectar guides, tubular-funnelform, 10–16 mm, glandular-pubescent externally, moderately yellow-pilose to yellow-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–5 mm diam., prominently 2-ridged abaxially; stamens: longer pair reaching orifice or slightly exserted, pollen-sacs opposite, navicular, 0.6–0.8 (–1) mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 6–9 mm, included, 0.3–0.6 mm diam., tip straight to recurved, glabrous or distal 1–3 mm sparsely to moderately pilose, hairs yellow, to 0.5 mm; style 9–12 mm. Capsules 4–6 × 2.5–3.5 mm, glabrous.

Discussion

Varieties 3 (3 in the flora).

Penstemon heterodoxus comprises three partially sympatric varieties centered in the Sierra Nevada of eastern California.

Selected References

None.

Key

1 Thyrse axes usually sparsely glandular-pubescent; n California. Penstemon heterodoxus var. shastensis
1 Thyrse axes moderately to densely glandular-pubescent; e California, wc Nevada. > 2
2 Stems 5–25 cm; basal and proximal cauline leaves 5–50 × 3–10 mm; verticillasters (1–)3; ec California, wc Nevada. Penstemon heterodoxus var. heterodoxus
2 Stems 15–35 cm; basal and proximal cauline leaves (17–)20–75 × 6–18 mm; verticillasters 2–6; se California. Penstemon heterodoxus var. cephalophorus

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon heterodoxus"
obtuse +  and acute +
Craig C. Freeman +
A. Gray in A. Gray et al. +
glandular-pubescent +
alternate +, subopposite +, whorled +  and opposite +
glaucescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
cauline +, proximal +  and basal +
rarely +, not glaucous +, glabrate +, pubescent +, hairy +, glandular-lanate +, glandular-pubescent +, puberulent +, glandular +  and glabrous +
not +  and leathery +
tapered;tapered +
oblong;oblanceolate or lanceolate +
not +  and leathery +
glandular-pubescent +
oblong;lanceolate +
0.7mm;1.2mm +
short-tubular +  and short-campanulate +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
sessile +  and short-petiolate +
6 cm60 mm <br />0.06 m <br /> (7 cm70 mm <br />0.07 m <br />) +
1 cm10 mm <br />0.01 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
petiolate +  and sessile +
Sierra beardtongue +
personate +, not personate +, bilabiate +  and symmetric +
lavender +  and blue violet or purple +
moderately yellow-pilose +  and yellow-lanate +
tubular-funnelform +
ampliate +
1 cm10 mm <br />0.01 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
2-6-flowered +
scale-like +  and not needlelike +
Calif. +  and Nev. +
drupe-like +
curved +  and straight +
1 +  and 4 +
0 cm0 mm <br />0 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
basal +  and cauline +
not leathery +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
glutinous +
glandular-ciliolate +, hairy +, puberulent +, glabrescent +, glabrous +  and glandular-pubescent +
lacerate +, toothed +  and entire +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
glandular-pubescent +
glandular-pubescent +
5 +  and 4 +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
navicular-sigmoid +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
1/5 +  and 2/3 +
0.4 cm4 mm <br />0.004 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Syn. Fl. N. Amer. +
minute +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
black +, gray +, brown +  and tan +
angled +  and slightly rounded +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (?) +  and 0.9 cm9 mm <br />0.009 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
erect;ascending;erect;ascending;decumbent +
not glaucous +, glandular-pubescent +, hairy +  and glabrous +
5 cm50 mm <br />0.05 m <br /> (65 cm650 mm <br />0.65 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
papillate +
denticulate +
Penstemon heterodoxus +
Penstemon sect. Penstemon +
species +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
2-ridged +  and inflated +
not constricted +
cylindric +
1 cm10 mm <br />0.01 m <br /> (12 cm120 mm <br />0.12 m <br />) +
straight +
pilose +  and glabrous +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
gibbous +  and not spurred +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
cauline +  and proximal +