Potentilla effusa var. effusa

Synonyms: Potentilla effusa var. filicaulis Nuttall P. hippiana var. filicaulis (Nuttall) B. Boivin
Treatment appears in FNA Volume 9. Treatment on page 163. Mentioned on page 164.

Leaflets gray to white, teeth (1–) 4–9 per side, abaxial surfaces moderately long-hairy especially on veins, moderately to densely cottony, adaxial moderately long-hairy and abundantly cottony. Flowers: epicalyx bractlets and sepals usually abundantly cottony at least proximally; anthers (0.6–) 0.7–1 mm. 2n = 56.


Phenology: Flowering summer.
Habitat: Dry rocky slopes, slightly moist meadows, grasslands, oak, aspen, and montane conifer woodlands (limestone pavement)
Elevation: 700–2600 m

Distribution

V9 237-distribution-map.jpg

Alta., Man., Sask., Colo., Idaho, Minn., Mont., Nebr., N.Mex., N.Dak., S.Dak., Utah, Wyo.

Discussion

Variety effusa occurs from southern Alberta to Manitoba southward on plains and in the mountains to central Colorado. Disjunct populations from northern foothills of the Uinta Mountains in Utah are included in var. effusa by B. C. Johnston (1980), but relegated to Potentilla hippiana by N. H. Holmgren (1997b). Verification is needed for single occurrences of the variety mapped by Johnston in eastern Idaho, northwestern Nebraska, and north-central New Mexico, without citation of specimens. A single collection from Valley County, Idaho, (E. A. Christenson 42, CONN) is almost certainly an introduction of unknown persistence.

The small, narrow, cottony-floccose epicalyx bractlets tend to be glabrescent at least distally, contrasting with the sharply acuminate sepals. Variety effusa, in its most distinctive form, has unevenly pinnate leaves with coarsely toothed gray leaflets that are equally cottony-floccose on both surfaces.

Collections of otherwise typical var. effusa from central Wyoming and adjacent Montana are uniformly smaller than average with only one to four teeth per side of the leaflets. These occur mainly on limestone outcrops (1300–2600 m elevation) and might qualify as a distinct variety.

Selected References

None.

Lower Taxa

None.

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white +  and more or less green +
cottony +  and long-hairy +
glabrescent +  and glabrate +
smooth +  and slightly rugose +
1.8mm +  and 2.1mm +
green +  and white +
cottony +  and long-hairy +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
acuminate +
Barbara Ertter +  and James L. Reveal +
Douglas ex Lehmann +
compound +  and simple +
15 cm150 mm <br />0.15 m <br /> (25 cm250 mm <br />0.25 m <br />) +
5 cm50 mm <br />0.05 m <br /> (15 cm150 mm <br />0.15 m <br />) +
Leucophyllae +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Alta. +, Man. +, Sask. +, Colo. +, Idaho +, Minn. +, Mont. +, Nebr. +, N.Mex. +, N.Dak. +, S.Dak. +, Utah +  and Wyo. +
not +  and aggregated +
700–2600 m +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
linear +  and narrowly lanceolate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
weak +  and stiff +
Dry rocky slopes, slightly moist meadows, grasslands, oak, aspen, and montane conifer woodlands (limestone pavement) +
stoloniferous +  and short-rhizomatous +
straight +
weak +  and stiff +
nearly +, abundant +  and sparse +
crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.5mm;2.5mm +
turbinate +, patelliform +  and cupulate +
crassinucellate +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
gray +  and white +
oblanceolate +  and narrowly obovate oblong or cuneate +
linear +  and oblanceolate +
palmate +  and pinnate +
Less (?) +, 1/4 (?) +  and 1/2 (?) +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
free +  and distinct +
4mm;6.5mm +
usually longer +
retuse;obcordate +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
5 cm50 mm <br />0.05 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
pinnate +  and subpinnate +
45 cm450 mm <br />0.45 m <br /> (50 cm500 mm <br />0.5 m <br />) +
3 cm30 mm <br />0.03 m <br /> (45 cm450 mm <br />0.45 m <br />) +
Nov. Stirp. Pug. +
not arillate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Endemic +  and Illustrated +
free +  and distinct +
not rooting +
reddish +  and green +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
deciduous +
distinct +
subapical +
papillate-swollen +  and tapered-filiform +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
gray +  and white or more or less green +
Potentilla effusa var. filicaulis +  and P. hippiana var. filicaulis +
not fleshy-thickened +
Potentilla effusa var. effusa +
Potentilla effusa +
variety +
4 (?) +  and 9 (?) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +