Agoseris heterophylla var. cryptopleura

Greene

Pittonia 2: 179. 1891.

Common names: California agoseris
Endemic
Synonyms: Agoseris californica (Nuttall) Hoover Agoseris heterophylla subsp. californica (Nuttall) Piper Agoseris heterophylla var. californica (Nuttall) Davidson & Moxley Agoseris heterophylla var. crenulata (H. M. Hall) Jepson Agoseris heterophylla var. turgida (H. M. Hall) Jepson Agoseris major Cryptopleura californica
Treatment appears in FNA Volume 19. Treatment on page 333. Mentioned on page 331, 332, 334.

Stems 0 or 1 (internodes to 1/2 times lengths of subtending leaves). Leaves erect to spreading; blades (2–) 5–15 (–24) cm × (1.5–) 2–10 (–16) mm, mostly toothed to lobed, rarely entire, teeth or lobes (2–) 3–5 pairs, faces usually densely, ± uniformly hairy, rarely glabrous. Peduncles 6–60 cm, lengths mostly 1.5–4 times leaves at flowering, 2–5 times length of leaves in fruit, glabrous or glabrate, or apically hairy to tomentose. Heads (7–) 10–18 mm wide (flowering). Phyllaries green or medially rosy purple, sometimes spotted, margins glabrous or ciliate, faces mostly hairy, occasionally glabrous or villous, stipitate-glandular, with translucent, yellowish or purple-septate, often purple-tipped hairs or eglandular with whitish-opaque hairs; outer erect to spreading, adaxially glabrous or pubescent and eglandular. Florets 20–100 (–300); corollas much surpassing phyllaries at flowering, tubes 2–5 mm, ligules 10–15 × 2–3 mm; anthers 2–4 mm. Cypselae: outermost strongly differing from inner, highly variable as to color, shape, ornamentation, and pubescence, ribs flattened to alate, or 0, straight to strongly undulate, often strongly diminishing proximally. 2n = 18.


Phenology: Flowering Mar–Sep.
Habitat: Mesic to dry habitats in grasslands, chaparral, oak woodlands, and open pine forests
Elevation: 150–2100 m

Discussion

Variety cryptopleura occurs on hills and ranges surrounding the Great Central Valley (it appears to be absent from the valley itself). It is ± sympatric with var. heterophylla, except that var. cryptopleura almost completely supplants var. heterophylla in the south Coast Ranges of California. Older floras treated this taxon as var. californica; more recent works have not recognized it at all. K. L. Chambers (1963b) demonstrated that var. cryptopleura is strictly allogamous and is not self-fertile. Varieties heterophylla and cryptopleura are differentiated almost entirely by corolla size, which appears to be correlated with breeding system; otherwise the two exhibit almost complete morphologic overlap. Corolla size in depauperate specimens of var. cryptopleura approaches that found in robust specimens of var. heterophylla. The two varieties still separate on corollas longer than the phyllaries in var. cryptopleura versus subequaling the phyllaries in var. heterophylla. The degree of introgression, if any, between var. cryptopleura and var. heterophylla is not known.

Selected References

None.

Lower Taxa

None.

"fine" is not a number."-5timeslengthofleaves" is not declared as a valid unit of measurement for this property. "-4timesleaves" is not declared as a valid unit of measurement for this property.

0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
scarious +
ovate +  and lanceolate +
corymbiform +  and paniculiform +
indeterminate +  and determinate +
Gary I. Baird +
Greene +
decurrent +
Macrorhynchus heterophyllus +
1-4 times bodies +
0.5 cm5 mm <br />0.005 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
compound +  and simple +
15 cm150 mm <br />0.15 m <br /> (24 cm240 mm <br />0.24 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
5 cm50 mm <br />0.05 m <br /> (15 cm150 mm <br />0.15 m <br />) +
mostly toothed;lobed +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
2mm;10mm +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
tuberculate +, rugose +, muricate +  and smooth +
tumid;mostly fusiform;obconic +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
filiform +
2 (?) +  and 6 (?) +
acute +, linear-lanceolate +  and cuneate-oblanceolate +
California agoseris +
pistillate +  and staminate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
papillate +
tapered +, beaked +, flattened +, obcompressed +, compressed +, prismatic +, fusiform +, ellipsoid +, columnar +  and clavate +
150–2100 m +
stipitate-glandular +, villous +, glabrous +  and hairy +
winged +  and beaked +
dispersed +
Mesic to dry habitats in grasslands, chaparral, oak woodlands, and open pine forests +
eglandular +, purple-tipped +  and purple-septate +
whitish-opaque +  and yellowish +
indeterminate +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
each +  and sessile +
pistillate +  and staminate +
cylindric;hemispheric +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
spreading;antrorse +
linear +  and spatulate +
hairy +  and glabrous +
ciliate +  and glabrous +
lobed +  and entire +
erect +  and spreading +
pubescent +  and glabrous +
2-carpellate +
pistillate +  and staminate +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
glabrate or apically hairy +  and tomentose +
6 cm60 mm <br />0.06 m <br /> (60 cm600 mm <br />0.6 m <br />) +
not purplish +
Flowering Mar–Sep. +
connate +  and distinct +
subequal +  and unequal +
bearing subulate enations +, hairy +  and bristly +
flat;convex +
straight +
undulate +  and flattened +
diminishing +
exalbuminous +
prostrate +  and ascending +
appendaged +  and truncate +
papillate +
Agoseris californica +, Agoseris heterophylla subsp. californica +, Agoseris heterophylla var. californica +, Agoseris heterophylla var. crenulata +, Agoseris heterophylla var. turgida +, Agoseris major +  and Cryptopleura californica +
Agoseris heterophylla var. cryptopleura +
Agoseris heterophylla +
variety +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +