Agoseris heterophylla var. heterophylla

Synonyms: Agoseris heterophylla var. glabra Suksdorf Agoseris heterophylla subsp. glabrata (Suksdorf) Piper Agoseris heterophylla subsp. normalis Piper Troximon heterophyllum var. cryptopleuroides
Treatment appears in FNA Volume 19. Treatment on page 333. Mentioned on page 334.

Stems 0 or 1 (internodes much shorter than subtending leaves). Leaves erect to spreading; blades (1–) 3–15 (–23) cm × (0.5–) 2–10 (–16) mm, mostly toothed, rarely lobed, lobes or teeth in 2–3 pairs, faces glabrous or densely, ± uniformly hairy. Peduncles 6–53 cm, lengths mostly (0.5–) 1.5–4.5 times leaves at flowering, (1.5–) 2–5 times leaves in fruit, glabrous or glabrate, or apically puberulent to villous. Heads 2–10 (–12) mm wide at flowering. Phyllaries green or medially rosy purple, sometimes spotted, margins glabrous or ciliate, faces mostly hairy, occasionally glabrous, or villous, stipitate-glandular, translucent, yellowish or purple-septate and often purple-tipped trichomes, or eglandular with whitish opaque trichomes; outer erect to spreading, adaxially glabrous or puberulent and eglandular. Florets (5–) 20–50 (–100); corollas ± equaling phyllaries at flowering, tubes 1–4 mm, ligules 2–6 × 1–2 mm; anthers 1–1.5 mm. Cypselae: outermost strongly differing from inner, highly variable in color, shape, ornamentation, and pubescence; ribs flattened to alate, or 0, straight to strongly undulate, not diminishing proximally. 2n = 18, 36.


Phenology: Flowering Mar–Sep.
Habitat: Wet to dry, mostly seasonal habitats in deserts, grasslands, chaparral, steppe, oak woodlands, and open pine forests, often disturbed sites
Elevation: 0–2300 m

Distribution

V19-512-distribution-map.gif

Introduced; B.C., Ariz., Calif., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Mexico (Baja California), Mexico (Guadalupe Island), Europe (Sweden)

Discussion

Variety heterophylla is the most widespread of the varieties. It exhibits remarkable variation in cypsela morphology. K. L. Chambers (1963b) demonstrated that this small-flowered variety is strongly autogamous and does not outcross. The same breeding system may be present in other Agoseris taxa that exhibit reduced corolla and anther size (e.g., A. apargioides var. maritima, A. grandiflora, A. aurantiaca). Hybridization does not appear to be common; apparent intermediates with A. apargioides and A. hirsuta are known.

Selected References

None.

Lower Taxa

None.

"fine" is not a number."(" is not declared as a valid unit of measurement for this property.

0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
scarious +
ovate +  and lanceolate +
corymbiform +  and paniculiform +
indeterminate +  and determinate +
Gary I. Baird +
(Nuttall) Greene +
decurrent +
Macrorhynchus heterophyllus +
1-4 times bodies +
0.5 cm5 mm <br />0.005 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
compound +  and simple +
15 cm150 mm <br />0.15 m <br /> (23 cm230 mm <br />0.23 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
3 cm30 mm <br />0.03 m <br /> (15 cm150 mm <br />0.15 m <br />) +
lobed;toothed +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
2mm;10mm +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
tuberculate +, rugose +, muricate +  and smooth +
tumid;mostly fusiform;obconic +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
filiform +
2 (?) +  and 6 (?) +
acute +, linear-lanceolate +  and cuneate-oblanceolate +
pistillate +  and staminate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
papillate +
tapered +, beaked +, flattened +, obcompressed +, compressed +, prismatic +, fusiform +, ellipsoid +, columnar +  and clavate +
B.C. +, Ariz. +, Calif. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +, Mexico (Baja California) +, Mexico (Guadalupe Island) +  and Europe (Sweden) +
0–2300 m +
eglandular +  and purple-septate +
stipitate-glandular +, villous +, glabrous +, hairy +  and densely +
winged +  and beaked +
dispersed +
Wet to dry, mostly seasonal habitats in deserts, grasslands, chaparral, steppe, oak woodlands, and open pine forests, often disturbed sites +
indeterminate +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
each +  and sessile +
pistillate +  and staminate +
cylindric;hemispheric +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
spreading;antrorse +
linear +  and spatulate +
hairy +  and glabrous +
ciliate +  and glabrous +
lobed +  and entire +
erect +  and spreading +
puberulent +  and glabrous +
2-carpellate +
pistillate +  and staminate +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
glabrate or apically puberulent +  and villous +
6 cm60 mm <br />0.06 m <br /> (53 cm530 mm <br />0.53 m <br />) +
not purplish +
Flowering Mar–Sep. +
connate +  and distinct +
subequal +  and unequal +
bearing subulate enations +, hairy +  and bristly +
flat;convex +
straight +
undulate +  and flattened +
diminishing +
exalbuminous +
prostrate +  and ascending +
appendaged +  and truncate +
papillate +
Agoseris heterophylla var. glabra +, Agoseris heterophylla subsp. glabrata +, Agoseris heterophylla subsp. normalis +  and Troximon heterophyllum var. cryptopleuroides +
Agoseris heterophylla var. heterophylla +
Agoseris heterophylla +
variety +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +