Alyssum simplex

Rudolphi

J. Bot. (Schrader) 1799(2): 290. 1799.

Synonyms: Alyssum parviflorum var. micranthum (C. A. Meyer) Dorn
Treatment appears in FNA Volume 7. Treatment on page 249. Mentioned on page 248, 250.

Annuals; canescent throughout, trichomes appressed, coarse, (3–) 5–10-rayed, rays sometimes unequal. Stems often several from base, usually erect or ascending, rarely decumbent, (unbranched or branched distally), (0.3–) 0.7–3 (–4) dm. Cauline leaves subsessile or (proximal) attenuate to petiolelike base (to 0.4 cm); blade oblanceolate, obovate-spatulate, or elliptic-lanceolate, (0.5–) 0.7–2.2 (–3) cm × (1–) 2–6 (–8) mm (gradually smaller distally), base attenuate, apex acute. Fruiting pedicels divaricate, straight, stout, (2–) 3–5 (–6) mm, trichomes stellate (some with unequal rays). Flowers: sepals oblong, 1.7–2.3 × 0.8–1.1 mm, trichomes stellate; petals pale-yellow, oblanceolate, (1.8–) 2–2.8 (–3.2) × 0.6–0.8 mm, base attenuate, apex often emarginate, sparsely stellate-pubescent abaxially or glabrous; filaments: median pairs broadly winged, apically 1-toothed or 2-toothed, lateral pair with broadly winged appendage, apically 2-cleft, 1.4–2 mm; anthers ovate, 0.3–0.4 mm. Fruits orbicular, (3.5–) 4–6.5 (–7) mm diam., apex truncate or shallowly emarginate; valves uniformly inflated at middle, broadly flattened at margins, densely stellate-pubescent (trichomes coarse, rays equal or unequal); ovules 2 per ovary; style 0.7–1.6 mm, basally stellate-pubescent or glabrous. Seeds ovoid, 1.6–2 × 1.2–1.5 mm, slightly compressed, margined or not. 2n = 16.


Phenology: Flowering Apr–Jun.
Habitat: Sagebrush areas, fields, roadsides, sandy slopes
Elevation: 1400-2400 m

Distribution

V7 321-distribution-map.gif

Introduced; Ariz., Calif., Colo., Kans., Mont., Nebr., Nev., N.Mex., Oreg., Utah, Wyo., Europe, c, sw Asia, n Africa

Discussion

Almost all European and North American authors, including T. R. Dudley (1964) and R. C. Rollins (1993), have used the name Alyssum minus (Linnaeus) Rothmaler for this species. Rothmaler made his new combination based on the nomen nudum Clypeola minus Linnaeus and cited A. campestre Linnaeus as a synonym. Therefore, that combination is illegitimate, and the species should be known as A. simplex, the earliest valid and legitimate name.

Most of the North American records of Alyssum strigosum are based on misidentified plants of A. simplex. The two species are similar in nearly all aspects except for trichome morphology of the fruit. In A. simplex, all fruit trichomes are stellate and occasionally some trichomes have rays coarser than the others. By contrast, the fruit trichomes in A. strigosum are markedly dimorphic and consist of stellate hairs mixed with tuberculate-based, much coarser, forked ones. I have not yet seen any material of A. strigosum from North America, and those annotated as such belong to A. simplex. T. R. Dudley (1968) and R. C. Rollins (1993) based their record of A. strigosum on a single collection, Davis 108 (DS), made in 1920 from Santa Clara County, California. Although I have not examined that specimen to verify its identity, it appears that the species has not become established in the flora area, and it is not included in the present account.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Alyssum simplex"
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
emarginate +  and truncate +
glabrous +  and stellate-pubescent +
Ihsan A. Al-Shehbaz +
Rudolphi +
sessile +  and petiolate +
not +  and rosulate +
attenuate +
appendaged +  and unappendaged +
2.2 cm22 mm <br />0.022 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
dissected +, repand +, sinuate +, crenate +  and dentate +
elliptic-lanceolate;obovate-spatulate;elliptic-lanceolate;obovate-spatulate;oblanceolate +
much smaller +  and reduced +
2mm;6mm +
urceolate +, campanulate +  and tubular +
branched +  and simple +
attenuate to petiolelike +  and subsessile +
well-developed +
distinct +
not +  and differentiated +
emarginate +  and entire +
incumbent +  and accumbent +
Ariz. +, Calif. +, Colo. +, Kans. +, Mont. +, Nebr. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wyo. +, Europe +, c +, sw Asia +  and n Africa +
1400-2400 m +
straight +  and curved +
connate +  and distinct +
lateral +  and median +
2-toothed +  and 1-toothed +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
actinomorphic +
latiseptate +  and sessile +
orbicular +
latiseptate +  and unsegmented +
Sagebrush areas, fields, roadsides, sandy slopes +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
rudimentary +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Flowering Apr–Jun. +
trinucleate +  and 3(-11)-colpate +
J. Bot. (Schrader) +
elongated +
reflexed +  and ascending +
stout +  and slender +
not +  and branched +
unequal +
not +  and mucilaginous +
not +  and margined +
aseriate +  and biseriate +
not +  and mucilaginous +
White (?) +, Black (?) +, Brown (?) +  and Yellow (?) +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
compressed +  and ovoid +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
persistent +  and caducous +
distinct +
erect +  and spreading +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
ascending +  and erect +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
glabrous +  and stellate-pubescent +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
Alyssum parviflorum var. micranthum +
Alyssum simplex +
species +
simple +  and sessile +
stellate-pubescent +
flattened +  and inflated +
anastomosing +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +