familyPottiaceae
subfamilyPottiaceae subfam. Barbuloideae
genusAnoectangium
speciesAnoectangium handelii

Anoectangium handelii

Schiffner

Ann. K. K. Naturhist. Hofmus. 27: 490, figs. 51–59. 1913,.

Richard H. Zander, Patricia M. Eckel
Treatment appears in FNA Volume 27. Treatment on page 522. Mentioned on page 521, 537, 538.
Please click on the illustration for a higher resolution version.
Illustrator: Patricia M. Eckel

Copyright: Flora of North America Association

Leaves dense, hiding the stem, ligulate to short-elliptic or ovate, 0.4–0.5 (–0.6) mm; apex broadly acute to rarely rounded, apiculus absent or of one cell; margins often 2-stratose in patches; costa usually ending (1–) 3–4 cells before apex. Specialized asexual reproduction by ovate gemmae, sometimes uncommon, occasionally apiculate by a projection, of ca. 8 cells, born on rhizoids in leaf-axils. Sporophytes unknown.


Habitat: Fissures in rock, calcareous and noncalcareous sandstone
Elevation: moderate to high elevations (900-1700 m)

Distribution

V27 751-distribution-map.gif

Calif., Colo., Nev., s Europe, sw, c Asia

Discussion

Anoectangium handelii is quite similar to Gymnostomum viridulum in habitat, size, general appearance, and production of axillary gemmae (a morphological phenocopy), but is immediately distinguished by its narrow adaxial costal groove. Though apparently rare (California in Inyo County, Colorado in Boulder and Larimer counties, Nevada in Clark County), its small size and similarity to sterile Gymnostomum make A. handelii easy for collectors to pass over. C. C. Heyn and I. Hernnstadt (2004b) provided additional discussion of this curious species.

Selected References

Zander, R. H. and W. A. Weber. 2005. Anoectangium handelii (Bryopsida, Pottiaceae) in the New World. Bryologist 108: 47–49.

Lower Taxa

None.
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Abaxial cell shape
elongate +
Abaxial epidermis fusion
distinct +
Abaxial epidermis variability
differentiated +
Adaxial cell shape
elongate +
Adaxial epidermis development
developed +
Adaxial epidermis presence
absent +
Adaxial outgrowth presence
absent +
Adaxial surface architecture
grooved +
Adaxial surface shape
keeled +
Annulus architecture
revoluble +
Annulus duration
deciduous +
Annulus variability
of 1-2 rows +  and differentiated +
Apex architecture
long-excurrent +  and short-excurrent +
Apex shape
broadly acute +  and rarely rounded +
Apiculu presence
absent +
Apiculu quantity
of one cell +
Author
Richard H. Zander +  and Patricia M. Eckel +
Authority
Schiffner +
Axillary hair coloration
brownish +  and hyaline +
Axillary hair position
basal +
Axillary hair quantity
1;2 +
Band fragility
strong +
Band presence
absent +
Band shape
semicircular +  and ovate +
Basal-cell architecture
thick-walled +
Basal-cell quantity
1;2;4 +
Basal-cell variability
differentiated +
Basal-cell width
wider +
Basal laminal cell pubescence or shape
bulging +
Basal laminal cell relief
papillose +  and smooth +
Basal laminal cell shape
Rectangular (?) +
Basal laminal cell variability
differentiated +
Basal laminal cell width
usually slightly wider +
Basal membrane architecture
trabeculate +
Basal membrane height
high +
Basal membrane position
low +
Basal membrane presence
absent +
Base shape
usually ovate;oblong +
Basionyms
Pottiaceae tribe Barbuleae +
Body shape
obovoid +
Branchlet height or length or size
short +
Calyptra architecture or pubescence or relief
smooth +
Calyptra atypical some measurement
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Calyptra relief
papillose +
Calyptra shape
cucullate +
Calyptra some measurement
0.12 cm1.2 mm <br />0.0012 m <br /> (?) +  and 0.15 cm1.5 mm <br />0.0015 m <br /> (?) +
Capsule atypical some measurement
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Capsule coloration
yellowbrown +  and brown +
Capsule dehiscence
cleistocarpous +  and stegocarpous +
Capsule shape
ovoid;elliptic +
Capsule some measurement
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Cell-wall development
specialized +
Cell-wall prominence or shape
flat +
Cell-wall pubescence or shape
bulging +
Cell-wall reproduction
dioicous +  and sexual +
Cell-wall shape
superficially flat +  and bulging +
Cell-wall size or width
thickened +
Cell-wall width
thin;greatly thickened +
Cell architecture
vesiculose +  and thin-walled +
Cell atypical quantity
1 (3) +
Cell coloration
brownish +
Cell quantity
8 +, 3 +  and 4 +
Cell shape
elongate-rhomboid +  and elongate +
Central scleroderm presence
absent +
Central strand fragility
strong +
Central strand presence
absent +
Central strand pubescence
tomentose +  and bare +
Costa architecture
short-excurrent +
Costa development
specialized +
Distal laminal cell architecture
2-stratose +
Distal laminal cell arrangement
stacked +
Distal laminal cell quantity
1 (?) +
Distal laminal cell shape
Rhomboid (?) +, Rarely (?) +, Hexagonal (?) +  and Subquadrate (?) +
Distal laminal cell size
small +
Distal laminal cell width
9mm;16mm +
Distal medial cell atypical quantity
1 (2) +
Distal medial cell quantity
1 +
Distal medial cell shape
elongate;subquadrate +
Distal medial cell width
7um (9um) +
Distribution
Calif. +, Colo. +, Nev. +, s Europe +, sw +  and c Asia +
Division architecture
spiculose +
Division shape
filiform +
Elevation
moderate to high elevations (900-1700 m) +
Epidermis variability
differentiated +
Exothecial cell shape
rectangular +
Gemma architecture
multicellular +
Gemma architecture or shape
apiculate +
Gemma presence
absent +
Gemma quantity
uncommon +
Gemma reproduction
asexual +
Gemma shape
Ovate (?) +
Group size
small +
Guide cell quantity
2 +  and 4 +
Habitat
Fissures in rock, calcareous and noncalcareous sandstone +
Hyalodermi presence
absent +
Hyalodermi quantity
rare +
Hydroid strand presence
absent +
Illustration
Present +
Illustration copyright
Flora of North America Association +
Illustrator
Patricia M. Eckel +
Interior perichaetial leaf architecture or shape
convolute-sheathing +
Interior perichaetial leaf shape
ovate-acuminate +
Interior perichaetial leaf some measurement
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Koh coloration
yellow +  and orange-red or red +
Koh quantity
2% +
Laminal cell shape
Rhomboidal (?) +
Laminal margin character
shape +
Laminal margin shape
plane +  and ovate +
Lateral branch height or length or size
short +
Layer atypical quantity
1 (3) +
Layer quantity
1 +
Leaf-stem arrangement or shape
contorted +
Leaf-stem distance
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Leaf-stem fixation or orientation
appressed +
Leaf-stem orientation
spreading +
Leaf-stem shape
concave;keeled;channeled;ovoid;lanceolate or lingulate +
Leaf architecture
ligulate +
Leaf arrangement
crowded +  and distant +
Leaf atypical some measurement
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Leaf base orientation
recurved +
Leaf base shape
denticulate +, crenulate +  and entire +
Leaf base variability
differentiated +
Leaf density
dense +
Leaf fragility
fragile +
Leaf orientation
spreading +  and appressed-incurved +
Leaf shape
ligulate-elliptic +  and short-elliptic +
Leaf size
reduced +
Leaf some measurement
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Margin architecture
2-stratose +
Margin architecture or shape
dentate +  and entire +
Margin orientation
recurved +
Margin shape
plane +
Margin shape or vernation
involute +
Neck presence
absent +
Neck size
small +
Number of lower taxa
0 +
Operculum atypical some measurement
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
Operculum shape
long-rostrate +
Operculum some measurement
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
Outgrowth nutrition
photosynthetic +
Papilla architecture
simple +
Papilla architecture or shape
multifurcating +
Papilla presence
absent +
Papilla shape
2-fid +
Papilla size
massive +
Perichaetial leaf architecture or shape
sheathing +
Perichaetial leaf shape
long-oval +
Perichaetial leaf size
usually larger +
Perichaetium position
lateral +  and terminal +
Perichaetium position or structure subtype
terminal +
Perigonium position
lateral +  and terminal +
Perigonium position or structure subtype
terminal +
Peristome presence
absent +
Peristome tooth architecture
twisted +
Peristome tooth arrangement or course or shape
linear +
Peristome tooth coloration
red +, orange +  and yellow +
Peristome tooth orientation
erect +
Peristome tooth presence
absent +
Peristome tooth prominence
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Peristome tooth quantity
32 +
Proximal strand quantity
1 (3) +
Publication title
Ann. K. K. Naturhist. Hofmus. +
Reference
zander2005a +
Row course
straight +
Row orientation or shape
oblique +
Row quantity
2;2;3 +
Scleroderm fragility
weak +
Scleroderm presence
absent +
Seta architecture
twisted +
Seta architecture or arrangement or growth form
solitary +
Seta coloration
yellowbrown +
Seta shape
elongate +
Seta some measurement
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Side quantity
2 +  and 1 +
Source xml
https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27 751.xml +
Spore atypical some measurement
12um (19um) +
Spore coloration
yellow +  and yellow-orange +
Spore relief
Papillose (?) +
Spore size
larger +
Spore some measurement
9um (12um) +
Stem architecture
branching +
Stem architecture or shape
sheathing +
Stem atypical some measurement
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Stem presence
absent +
Stem shape
oval +  and rounded-triangular or pentagonal +
Stem some measurement
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Taxon family
Pottiaceae +
Taxon name
Anoectangium handelii +
Taxon parent
Anoectangium +
Taxon rank
species +
Theca shape
ovoid;cylindric +
Tomentum coloration
redbrown +
Transverse-section shape
Oval (Reniform) +
Volume
Volume 27 +
Wall coloration
red +  and yellow +
Wall relief
Papillose (?) +  and Smooth (?) +
Wall width
thin +
Whole-organism shape
acuminate +, triangular +  and lanceolate +
Whole organism architecture
stemless +  and perigoniate +
Whole organism architecture or shape
budlike +
Whole organism coloration
green;yellowbrown distally +
Whole organism growth form
plant +
Whole organism size
occasionally smaller than the perichaetiate +