Anoectangium handelii

Schiffner

Ann. K. K. Naturhist. Hofmus. 27: 490, figs. 51–59. 1913,.

Treatment appears in FNA Volume 27. Treatment on page 522. Mentioned on page 521, 537, 538.

Leaves dense, hiding the stem, ligulate to short-elliptic or ovate, 0.4–0.5 (–0.6) mm; apex broadly acute to rarely rounded, apiculus absent or of one cell; margins often 2-stratose in patches; costa usually ending (1–) 3–4 cells before apex. Specialized asexual reproduction by ovate gemmae, sometimes uncommon, occasionally apiculate by a projection, of ca. 8 cells, born on rhizoids in leaf-axils. Sporophytes unknown.


Habitat: Fissures in rock, calcareous and noncalcareous sandstone
Elevation: moderate to high elevations (900-1700 m)

Distribution

V27 751-distribution-map.gif

Calif., Colo., Nev., s Europe, sw, c Asia

Discussion

Anoectangium handelii is quite similar to Gymnostomum viridulum in habitat, size, general appearance, and production of axillary gemmae (a morphological phenocopy), but is immediately distinguished by its narrow adaxial costal groove. Though apparently rare (California in Inyo County, Colorado in Boulder and Larimer counties, Nevada in Clark County), its small size and similarity to sterile Gymnostomum make A. handelii easy for collectors to pass over. C. C. Heyn and I. Hernnstadt (2004b) provided additional discussion of this curious species.

Lower Taxa

None.
... more about "Anoectangium handelii"
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
of one cell +
Richard H. Zander +  and Patricia M. Eckel +
Schiffner +
brownish +  and hyaline +
semicircular +  and ovate +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
Pottiaceae tribe Barbuleae +
obovoid +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
cucullate +
0.12 cm1.2 mm <br />0.0012 m <br /> (?) +  and 0.15 cm1.5 mm <br />0.0015 m <br /> (?) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
yellowbrown +  and brown +
cleistocarpous +  and stegocarpous +
ovoid;elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
dioicous +  and sexual +
superficially flat +  and bulging +
thin;greatly thickened +
vesiculose +  and thin-walled +
8 (?) +, 3 (?) +  and 4 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
short-excurrent +
specialized +
elongate;subquadrate +
Calif. +, Colo. +, Nev. +, s Europe +, sw +  and c Asia +
moderate to high elevations (900-1700 m) +
differentiated +
multicellular +
Fissures in rock, calcareous and noncalcareous sandstone +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
yellow +  and orange-red or red +
plane +  and ovate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
crowded +  and distant +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
denticulate +, crenulate +  and entire +
differentiated +
ligulate-elliptic +  and short-elliptic +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
dentate +  and entire +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
long-rostrate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
photosynthetic +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Ann. K. K. Naturhist. Hofmus. +
zander2005a +
straight +
yellowbrown +
elongate +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
2 +  and 1 +
yellow +  and yellow-orange +
oval +  and rounded-triangular or pentagonal +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Anoectangium handelii +
Anoectangium +
species +
ovoid;cylindric +
red +  and yellow +
acuminate +, triangular +  and lanceolate +
stemless +  and perigoniate +
green;yellowbrown distally +
occasionally smaller than the perichaetiate +