Antennaria howellii

Greene

Pittonia 3: 174. 1897.

Common names: Howell’s pussytoes antennaire de Howell
Endemic
Synonyms: Antennaria neglecta subsp. howellii (Greene) Hultén A. neglecta var. howellii (Greene) Cronquist A. neodioica subsp. howellii (Greene) R. J. Bayer
Treatment appears in FNA Volume 19. Treatment on page 403. Mentioned on page 390, 395, 401, 402, 404, 405, 406.

Gynoecious (staminate plants very uncommon). Plants (6–) 8–35 cm (stems sometimes stipitate-glandular). Stolons 1–9 (–12) cm. Basal leaves 1-nerved, spatulate to oblanceolate, spatulate-obovate, narrowly to broadly ovate, or cuneate-oblanceolate, 20–48 (–65) × 2.5–20 mm, tips mucronate, faces abaxially tomentose, adaxially green-glabrous or gray-pubescent. Cauline leaves linear, 8–40 mm, distal sometimes flagged (apices acute). Heads 3–15 in corymbiform arrays. Involucres: staminate 6–6.5 mm; pistillate 6–11 mm. Phyllaries (bases sometimes rose) distally white, cream, or light-brown. Corollas: staminate 3–4 mm; pistillate 3.5–6.5 (–8) mm. Cypselae 0.8–2 mm, ± papillate; pappi: staminate 4–4.5 mm; pistillate 5.5–9 mm. 2n = 56, 84, 140 (under A. neodioica).

Distribution

V19-648-distribution-map.gif

Alta., B.C., Man., N.B., N.S., N.W.T., Nfld. and Labr. (Nfld.), Ont., P.E.I., Que., Sask., Yukon, Calif., Colo., Conn., Del., Idaho, Ill., Ind., Iowa, Kans., Ky., Maine, Mass., Md., Mich., Minn., Mont., N.C., N.Dak., N.H., N.J., N.Y., Nebr., Ohio, Oreg., Pa., R.I., S.Dak., Tenn., Utah, Va., Vt., W.Va., Wash., Wis., Wyo.

Discussion

Subspecies 4 (4 in the flora).

The Antennaria howellii (previously A. neodioica) polyploid complex is highly variable morphologically; four more or less distinct subspecies can be recognized within it. The sexual progenitors of the complex are A. neglecta, A. plantaginifolia, A. racemosa, and A. virginica (see R. J. Bayer 1985). Antennaria marginata may also be a minor contributor to the origins of the complex. A. Cronquist (H. A. Gleason and Cronquist 1991) included members of this complex in A. neglecta; I maintain, because these apomicts are of hybrid polyploid origin from among multiple sexual progenitors, they best not be included within the circumscription of any one sexual progenitor (Bayer 1989d).

Selected References

None.

Key

1 Basal leaves green-glabrous adaxially > 2
1 Basal leaves pubescent adaxially (sometimes glabrescent with age) > 3
2 Basal leaves 1-nerved, 20–40 × 6–9 mm; distal cauline leaves flagged; phyllaries distally white or cream Antennaria howellii subsp. canadensis
2 Basal leaves 1–3-nerved, 25–40 × 9–12 mm; distal cauline leaves not flagged; phyllaries distally light brown or white Antennaria howellii subsp. howellii
3 Basal leaves spatulate to narrowly or broadly obovate (petiolate); stolons 3–8(–12) cm (leaves along stolons almost equal to those in rosettes at ends) Antennaria howellii subsp. neodioica
3 Basal leaves cuneate-oblanceolate, spatulate, or spatulate-obovate (without distinct petioles); stolons 4–9 cm (leaves along stolons smaller than in rosettes at ends) Antennaria howellii subsp. petaloidea
... more about "Antennaria howellii"
140 +, 84 +  and 56 +
more or less tomentose +  and sericeous or glabrescent +
scarious +
mostly ovate +  and lanceovate or linear +
indeterminate +  and determinate +
Randall J. Bayer +
Greene +
4.8 cm48 mm <br />0.048 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
2 cm20 mm <br />0.02 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (2 cm20 mm <br />0.02 m <br />) +
decurrent +
compound +  and simple +
spatulate;oblanceolate;spatulate;oblanceolate;linear;lanceolate;elliptic;cuneate +
5-ribbed +, 3-ribbed +  and 2-ribbed +
muriculate +, papillate +  and smooth +
0.8 cm8 mm <br />0.008 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Howell’s pussytoes +  and antennaire de Howell +
pistillate +  and staminate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
red +, yellow +  and white +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
papillate +
mostly ellipsoid +  and ovoid +
pistillate +  and staminate +
Alta. +, B.C. +, Man. +, N.B. +, N.S. +, N.W.T. +, Nfld. and Labr. (Nfld.) +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Calif. +, Colo. +, Conn. +, Del. +, Idaho +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Maine +, Mass. +, Md. +, Mich. +, Minn. +, Mont. +, N.C. +, N.Dak. +, N.H. +, N.J. +, N.Y. +, Nebr. +, Ohio +, Oreg. +, Pa. +, R.I. +, S.Dak. +, Tenn. +, Utah +, Va. +, Vt. +, W.Va. +, Wash. +, Wis. +  and Wyo. +
gray-pubescent +, green-glabrous +  and tomentose +
papillate +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
in corymbiform , paniculiform , racemiform or subcapitate , arrays +  and singly +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
pistillate +  and staminate +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
campanulate;cylindric +
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
rarely lanceovate +  and lanceolate +
2-carpellate +
pistillate +  and staminate +
persistent +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
bearing subulate enations +, hairy +  and bristly +
flat;convex or ovoid +
exalbuminous +
appendaged +  and truncate +
papillate +
Antennaria neglecta subsp. howellii +, A. neglecta var. howellii +  and A. neodioica subsp. howellii +
Antennaria howellii +
Antennaria +
species +
mucronate +
25 cm250 mm <br />0.25 m <br /> (70 cm700 mm <br />0.7 m <br />) +
4 cm40 mm <br />0.04 m <br /> (25 cm250 mm <br />0.25 m <br />) +